This is the page of Jesús Julio (Chechu) Camarero Martínez team. I am a plant-forest ecologist and dendrochronologist working on growth , regeneration, dieback and mortality processes in woody plants and, particularly, in trees. We are a research group using dendrochronology, i.e. the study of tree rings in woody plants, to solve ecological questions. We are interested in deciphering the causes and consequences of drought-induced forest decline. We apply retrospective approaches for inferring how forests are responding to past and current global changes and to reconstruct past climate (dendroclimatology). We study cambial biology and xylogenesis to understand how trees form wood and how climate may affect the pace and pattern of xylem growth.

I work in the INSTITUTO PIRENAICO DE ECOLOGÍA (CSIC) as researcher.


January 1976


Zaragoza, Spain

Bachelor’s Degree

Universidad de Zaragoza

Master’s Degree

Universidad de Barcelona


Xylem project

Cambial biology and xylogenesis.

In Xylogenesis and the functional xylem anatomy of woody plants
Dendro Chechu

Reconstructing climate and forest dynamics from tree-ring proxies.

In Mortality and dynamics of Pyrenean forests
Logging road

Growth in tropical dry forests.

In Retrospective growth analyses in tropical dry forests based on dendrochronology
Tesso project

Dencroecology in high elevation forests.

In Dynamics and dieback of Pyrenean forests and treelines


Point Pattern Analyses 

(introductory talk in Spanish).

Talk on forest decline (1). 

Dept. Ecology, Univ. Barcelona (in Spanish).

Talk on forest decline (2)

Dept. Ecology, Univ. Barcelona (in Spanish).

Forests and global change. 

A talk on the effects of global change of forests (in Spanish).

Consequences of forest decline for water use in trees. 

This is an example on how declining Pyrenean silver fir forests are also decreasing their water use efficient (the water they use for uptaking carbon). Links to the comment (in Spanish) and to the paper.

Alpine treelines.

A summary about our studies on treeline dynamics in the Pyrenees (in Spanish).

My life as dendrochronologist. 

Something about the way dendrochronologists live and love science (in Spanish).

Urban trees are living trees. 

I was always interested on urban trees and how they withstand urban life (in Spanish).


Current PhD Students.


Xavier Serra Maluquer

He evaluated how growth resilience to drought depends on forest site conditions, climate and functional traits (co-supervised with Dr. Antonio Gazol Burgos).


Cristina Valeriano Peñas

She is using tree rings and growth models to better understand how trees respond to climate (co-supervised with Dr. Raúl Sánchez Salguero).

Former PhD Students.


Jorge Albuixech

He studied the relationships between growth and masting in Mediterranean oak species (co-supervised with Dr. Gabriel Montserrat-Martí).


Arben Q. Alla

He compared primary and secondary growth in Mediterranean oak species (co-supervised with Dr. Gabriel Montserrat-Martí).


Edmond Pasho

He quantified the effects of drought on vegetation activity and radial growth (co-supervised with Dr. Sergio Vicente-Serrano).


J. Diego Galván

He analyzed how climate change affects growth in Pinus uncinata high-elevation forests and how this affects tree-ring proxies (co-supervised with Dr. Emilia Gutiérrez).


Hooz Angela Mendivelso

She analyed growth dynamics and demography in a tropical dry forest (co-supervised with Dr. Pieter Zuidema).


Gabriel Sangüesa Barreda

He studied how biotic stressors (mistletoe, defoliators, beetles) drive tree growth and function (co-supervised with Dr. Juan Carlos Linares).

Current PhD Students.


Gabriel Sangüesa Barreda

Studies how biotic stressors (mistletoe, defoliators, beetles) drive tree growth and function (co-supervised with Dr. J. Carlos Linares).


Hooz Angela Mendivelso

Is analyzing growth dynamics and demography in a tropical dry forest (co-supervised with Dr. Pieter Zuidema).


Xavier Serra Maluquer

Evaluated how growth resilience to drought depends on forest site conditions, climate and functional traits.

Former PhD Students.


Jorge Albuixech

Studied the relationships between growth and masting in Mediterranean oak species (co-supervised with Dr. Gabriel Montserrat-Martí).


Arben Q. Alla

Worked on primary and secondary growth in Mediterranean oak species (co-supervised with Dr. Gabriel Montserrat-Martí).


Edmond Pasho

Quantified the effects of drought on vegetation activity and radial growth (co-supervised with Dr. Sergio Vicente-Serrano).


J. Diego Galván

Analyzed how climate change affects growth in Pinus uncinata high-elevation forests and how this affects tree-ring proxies (co-supervised with Dr. Emilia Gutiérrez).


  • Jonàs Oliva is a forest pathologist at the Swedish University of Agricultural Sciences.
  • Eryuan Liang is dendrochronologist and dendroecologist at the Chinese Academy of Sciences.
  • Antonio Gazol is a postdoctoral researcher working on plant community patterns at the IPE (CSIC).
  • Juan Carlos Linares is an associate professor of ecology at Universidad Pablo de Olavide, Seville, Spain.
  • Emilia Gutiérrez is a professor of ecology at Universitat de Barcelona, Barcelona, Spain.
  • Thorsten Wiegand is a Senior Scientist and serves a ecologist at the Helmholtz Centre for Environmental Research, Leipzig, Germany.
  • Enric Batllori is an associate professor in Barcelona with who I enjoy working on treeline dynamics.
  • Pieter Zuidema is an associate professor on Forest Ecology at Wageningen University, The Netherlands.
  • Jacques Tardif is professor of ecology at University of Winnipeg, Canada.
  • Bosco Imbert is an associate professor of forest ecology at Universidad Pública de Navarra, Pamplona, Spain.
  • Lluís Coll is a forest ecologist working at the Centre Tecnològic Forestal de Catalunya, Solsona, Spain.
  • Josep Maria Espelta is a researcher at Centre de Recerca Ecològica i Aplicacions Forestals, Barcelona, Spain.
  • Alex Fajardo is a forest ecologist at Research Center of Patagonian Ecosystems, Coyhaique, Chile.


By obtaining articles from this page, you are agreeing to download no more than a single copy of any publication available at this site, to use these publications only for your own personal non-commercial purposes, and not to distribute copies to others or make copies available for distribution.


  1. Camarero, J.J., Colangelo, M., Rodríguez-González, P.M., Sánchez-Miranda, A., Sánchez-Salguero, R., Campelo, F., Rita, A. and Ripullone, F. 2021. Wood anatomy and tree growth covary in Mediterranean ash forests. Dendrochronologia (in press).
  2. Camarero, J.J., Gazol, A., Sánchez-Salguero, R., Fajardo, A., McIntire, E.J.B. and Liang, E. . 2021. Tree growth and treeline responses to temperature: Different questions and concepts. Global Change Biology DOI 10.1111/gcb.15728
  3. Puchi, P., Camarero, J.J., Battipaglia, G. and Carrer, M. 2021. Retrospective analysis of wood anatomical traits and tree-ring isotopes suggests site-specific mechanisms triggering Araucaria araucana drought-induced dieback. Global Change Biology DOI 10.1111/gcb.15881 PDF
  4. López, R., Cano, F.J., Rodríguez Calcerrada, J., Sangüesa-Barreda, G., Gazol, A., Camarero, J.J., Rozenberg, P. and Gil, L. 2021. Tree-ring density and carbon isotope composition are early-warning signals of drought-induced mortality in the drought tolerant Canary Island pine. Agricultural and Forest Meteorology (in press).
  5. Tamudo, E., Camarero, J.J., Sangüesa-Barreda, G. and Anadón, J.D. 2021. Dwarf mistletoe and drought contribute to growth decline, dieback and mortality of junipers. Forests (in press).
  6. González de Andrés, E., Suárez, M.L., Querejeta, J.I. and Camarero, J.J. 2021. Chronically low nutrient concentrations in tree rings are linked to greater tree vulnerability to drought in Nothofagus dombeyi. Forests (in press).
  7. Valeriano, C., Gazol, A., Colangelo, M., González de Andrés, E. and Camarero, J.J.  2021. Modeling climate impacts on tree growth to assess tree vulnerability to drought during forest dieback. Frontiers in Plant Science 10.3389/fpls.2021.672855 www PDF
  8. Wang, Y., Bo, L., and Camarero, J.J. 2021. Moisture mediates temperature-growth couplings of high-elevation shrubs in the Tibetan Plateau. Trees (in press).
  9. Dorigo, W.., Camarero, J.J. …2021. The International Soil Moisture Network: serving Earth system science for over a decade. Hydrology and Earth System Sciences (in press).
  10. Qiu, T., Aravena, M.-C., Andrus, R., Ascoli, D., Berberon, Y.., Camarero, J.J. ,…and Clark, J. S. 2021. Is there tree senescence? The fecundity evidence. Proceedings of the National Academy of Sciences of the United States of America 118: e2106130118. doi 10.1073/pnas.2106130118
  11. Colangelo, M., Camarero, J.J. , Gazol, A., Piovesan, G., Borghetti, M., Baliva, M., Gentilesca, T., Rita, A., Schettino, A. and Ripullone, F. 2021. Mediterranean old-growth forests exhibit resistance to climate warming. Science of the Total Environment 801: 149684 PDF
  12. Camarero, J.J. , Rubio-Cuadrado, A. and Gazol, A. 2021. Climate windows of intra-annual growth and post-drought recovery in Mediterranean trees. Agricultural and Forest Meteorology 308-309: 108606 PDF
  13. Shestakova, T.A., Camarero, J.J. and Voltas, J. 2021. Are global forests performing in sync? The need to account for spatiotemporal biases in tree-ring records. Journal of Biogeography (in press).
  14. Wang, Y., Liang, E., Lu, X., Camarero, J.J. , Babst, F., Shen, M. and Peñuelas, J. 2021. Warming-induced shrubline advance stalled by moisture limitation on the Tibetan Plateau. Ecography DOI 10.1111/ecog.05845 PDF
  15. Tumajer, J., Buras, A., Camarero, J.J. , Carrer, M., Shetti, R., Wilmking, M., Altman, J., Sangüesa-Barreda, G. and Lehejček, J. 2021. Growing faster, longer or both? Modelling plastic response of Juniperus communis growth phenology to climate change. Global Ecology and Biogeography doi 10.1111/geb.13377 PDF
  16. Camarero, J.J.  2021. The drought-dieback-death conundrum in trees and forests. Plant Ecology & Diversity (in press) DOI 10.1080/17550874.2021.1961172.
  17. Manrique-Alba, A., Beguería, A., Tomas-Burguera, M. and Camarero, J.J.  2021. Increased post-drought growth after thinning in Pinus nigra plantations. Forests 985: 1281062 PDF
  18. Camarero, J.J. , Gazol, A., Linares, J.C., Fajardo, A., Colangelo, M., Valeriano, C., Sánchez-Salguero, R., Sangüesa-Barreda, G., Granda, E. and Gimeno, T.E. 2021. Differences in temperature sensitivity and drought recovery between natural stands and plantations of conifers are species-specific. Science of the Total Environment 796: 148930 PDF
  19. González de Andrés, E., Rosas, T., Camarero, J.J. and Martínez-Vilalta, J. 2021. The intraspecific variation of functional traits modulates drought resilience of European beech and pubescent oak. Journal of Ecology (in press).
  20. Pompa-García, M., Camarero, J.J. , Colangelo, M. and González-Cásares, M. 2021. Inter and intra-annual links between climate, tree growth and NDVI: improving the resolution of drought proxies in conifer forests. International Journal of Biometeorology doi 10.1007/s00484-021-02170-5. PDF
  21. Camarero, J.J. , Colangelo, M., Gazol, A., Pizarro, M., Valeriano, C. and Igual, J.M. 2021. Effects of windthrows on forest cover, tree growth and soils in drought-prone pine plantations. Forests 12: 817 www PDF
  22. Moreno-Fernández, D., Viana-Soto, A., Camarero, J.J. , Zavala, M.A., Tijerín, J. and García, M. 2021. Using spectral indices as early warning signals of forest dieback: The case of drought-prone Pinus pinaster forests. Science of the Total Environment 793: 148578 PDF
  23. Sigdel, S.R., Pandey, J., Liang, E., Muhammad, S., Babst, F., Leavitt, S.W., Shen, M., Zhu, H., Salerno, F., Piao, S., Camarero, J.J. and Peñuelas, J. 2021. No benefits from warming even for subnival vegetation in the central Himalayas. Science Bulletin (in press), doi: 10.1016/j.scib.2021.06.005.
  24. Camarero, J.J. , Gazol, A., Sánchez-Salguero, R., Fajardo, A., McIntire, E.J.B. and Liang, E. 2021. Tree growth and treeline responses to temperature: different questions and concepts. Global Change Biology (in press), doi: 10.1111/gcb.15728.
  25. Garcia-Barreda, S., Sangüesa-Barreda, G., Madrigal-González, J., Seijo, F., González de Andrés, E. and Camarero, J.J. 2021. Reproductive phenology determines the linkages between radial growth, fruit production and climate in four Mediterranean tree species. Agricultural and Forest Meteorology 307: 108493. PDF
  26. Rubio-Cuadrado, A., Camarero, J.J. , Rodríguez-Calcerrada, J., Perea, R., Gómez, C.,
    Montes, F. and Gil, L. 2021. Impact of successive spring frosts on leaf phenology and radial growth in three deciduous tree species with contrasting climate requirements in central Spain. Tree Physiology (in press).
  27. Camarero, J.J. , Sangüesa-Barreda, G., Montiel-Molina, C., Luelmo-Lautenschlaeger, R., Ortega, P., Génova, M. and López-Sáez, J.A. 2021. Historical fires induced deforestation in relict Scots pine forests during the late 19th century. Fire 4: 29 www PDF
  28. Lucas-Borja, M.E., Andivia, E., Candel-Pérez, D., Linares, J.C. and Camarero, J.J. 2021. Long term forest management drives drought resilience in Mediterranean black pine forest. Trees (in press).
  29. García-Cervigón, A.I., Natalia Mercado, L., Mendivelso, H.A., Toledo, M. and Camarero, J.J. 2021. Adjusting xylem anatomy and growth to inter-annual climate variability in two Fabaceae species (Centrolobium microchaete, Cenostigma pluviosum) from Bolivian dry tropical forests. Dendrochronologia 67: 125840. PDF
  30. Marqués, L.,  Peltier, D.M.P., Camarero, J.J. , Zavala, M.A., Madrigal-González, J., Sangüesa-Barreda, G. and Ogle, K. 2021. Disentangling the legacies of climate and management on tree growth. Ecosystems doi 10.1007/s10021-021-00650-8. PDF
  31. Bose, A.K., Scherrer, D., Camarero, J.J. et al. 2021. Climate sensitivity and drought seasonality determine post-drought growth recovery of Quercus petraea and Quercus robur in Europe. Science of the Total Environment 784: 147222. PDF
  32. Lu, X., Camarero, J.J. and Liang, E. 2021. Threshold responses of juniper tree growth and regeneration to climate warming and drought stress at alpine treeline. Trees doi 10.1007/s00468-021-02135-6. PDF
  33. Zheng, T., Martínez-Vilalta, J., García-Valdés, R., Gazol, A., Camarero, J.J. and Mencuccini, M. 2021. Disentangling biology from mathematical necessity in twentieth-century gymnosperm resilience trends. Nature Ecology and Evolution www
  34. Camarero, J.J. , Valeriano, C., Gazol, A., Colangelo, M. and Sánchez-Salguero, R. 2021. Climate differently impacts growth of coexisting trees and shrubs under semi-arid Mediterranean conditions. Forests 12: 381 www PDF
  35. Camarero, J.J. , Gazol, A., Sangüesa-Barreda, G., Vergarechea, M., Alfaro-Sánchez, R., Cattaneo, N. and Vicente-Serrano, S.M. 2021. Tree growth is more limited by drought in rear-edge forests most of the times. Forest Ecosystems 8: 25 www PDF
  36. Rodríguez-González, P.M., Colangelo, M., Sánchez-Miranda, A., Sánchez-Salguero, R., Campelo, F., Rita, A., Gomes Marques, I., Albuquerque, A., Ripullone, F. and Camarero, J.J. 2021. Climate, drought and hydrology drive narrow-leaved ash growth dynamics in southern European riparian forests. Forest Ecology and Management 490: 119128. PDF
  37. Marqués, L., Camarero, J.J. , Zavala, M.A., Stoffel, M., Ballesteros-Cánovas, J.A., Sancho-García, C. and Madrigal-González, J. 2021. Evaluating tree-to-tree competition during stand development in a relict Scots pine forest: how much does climate matter? Trees doi 10.1007/s00468-021-02109-8. PDF
  38. Bai, M., Yao, Q., Camarero, J.J. , Hu, H., Dong, Z., Li, Y., Zhou, F., Chen, X., Guo, G., Cao, X. and Fang, F. 2020. El Niño-Southern Oscillation modulates insect outbreaks in humid subtropical China: evidences from tree rings and carbon isotopes. Dendrochronologia (in press).
  39. Pompa-García, M., Camarero, J.J. , Colangelo, M. and Gallardo-Salazar, J.L. 2021. Xylogenesis is uncoupled from forest productivity. Trees 35: 1123–1134. PDF
  40. Camarero, J.J. , Gazol, A., Sánchez-Salguero, R., Fajardo, A., McIntire, E.J.B., Gutiérrez, E., Batllori, E. et al. 2021. Global fading of the temperature-growth coupling at alpine and polar treelines. Global Change Biology doi 10.1111/gcb.15530. PDF
  41. Camarero, J.J. , Colangelo, M., Gazol A. and Azorín-Molina, C. 2020. Drought and cold
    spells trigger dieback in temperate forests. Dendrochronologia 66: 125802. PDF
  42. Pompa-García, M., González-Casares, M., Gazol, A. and Camarero, J.J.  2021. Run to the hills: forest growth responsiveness to drought increased at higher elevation during the late 20th century. Science of the Total Environment 772: 145286. PDF
  43. Camarero, J.J. , Colangelo, M., Gracia-Balaga, A., Ortega-Martínez, M.A. and Büntgen, U. 2021. Demystifying the age of old olive trees. Dendrochronologia 65: 125802. PDF


  1. Vicente-Serrano, S.M., Peña-Angulo, D., Murphy, C., López-Moreno, J.I., Tomàs-Burguera, M., Domínguez-Castro, F., Tian, F., Eklundh, L., Cai, Z., Álvarez-Farizo, B., Noguera, I., Camarero, J.J., Sánchez-Salguero, R., Gazol, A., Grainger, S., Conradt, T., Boincean, B. and El Kenawy, A. 2020. The complex multi-sectoral impacts of drought: Evidence from a mountainous basin in the Central Spanish Pyrenees. Science of the Total Environment 769: 144702. PDF
  2. Huang, J.-G., Ma, Q., Biondi, F., Deslauriers, A., Fonti, P., Liang, E., Mäkinen, H., Oberhuber, W., Rathgeber, C.B.K., Tognetti, R., Treml, V., Yang, B., Antonucci, S., Bergeron, Y., Camarero, J.J., Campelo, F., Čufar, K., Cuny, H.E., De Luis, M., Giovannelli, A., Gričar, J., Gruber, A., Gryc, V., Güney, A., Guo, X., Huang, W., Ivanova, A., Jyske, T., Kašpar, K., King, G., Krause, C., Lombardi, F., Martinez, E., Morin, H., Nöjd, P., Peters, R., Prislan, P., Saracino, A., Swidrak, I., Vavrčík, H., Vieira, J., Yu, B., Zhai, L., Zhang, S., Zeng, Q., Ziaco, E. and Rossi, S. 2020. Reply to Elmendorf and Ettinger: Photoperiod plays a dominant and irreplaceable role in triggering secondary growth resumption. Proceedings of the National Academy of Sciences of the United States of America 117: 32865-32867. doi 10.1073/pnas.2019931117 PDF
  3. Camarero, J.J. 2020. Within- versus between-species size effects on drought-induced dieback and mortality. Tree Physiology (in press).
  4. González de Andrés, E. and Camarero, J.J. 2020. Disentangling mechanisms of drought-induced dieback in Pinus nigra from growth and wood isotope patterns. Forests doi: 10.3390/f11121339. www PDF
  5. Serra-Maluquer, X., Granda, E., Camarero, J.J., Vilà-Cabrera, A., Jump, A., Sánchez-Salguero, R., Sangüesa-Barreda, G., Imbert, B. and Gazol, A. 2020. Impacts of recurrent dry and wet years alter long-term tree growth trajectories. Journal of Ecology doi: 10.1111/1365-2745.13579. PDF
  6. Shestakova, T.A., Mutke, S., Gordo, J., Camarero, J.J., Sin, E., Pemán, J. and Voltas, J. 2020. Weather as main driver for masting and stem growth variation in Stone pine supports compatible timber and nut co-production. Agricultural and Forest Meteorology 298-299: 108287. PDF
  7. Lu, X., Liang, E., Camarero, J.J. and Ellison, A.M. 2020. An unusually high shrubline on the Tibetan Plateau. Ecology – The Scientific Naturalist (in press).
  8. Gazol, A., Camarero, J.J., Sangüesa-Barreda, G., Serra-Maluquer, X., Sánchez-Salguero, R., Coll, Ll. and Casals, P. 2020.Tree species are differently impacted by cumulative drought stress and present higher growth synchrony in dry places. Frontiers in Forests and Global Change doi: 10.3389/ffgc.2020.573346 www PDF
  9. Camarero, J.J. and Rubio-Cuadrado, A. 2020. Relating climate, drought and radial growth in broadleaf Mediterranean tree and shrub species: new approaches to quantify climate-growth
    relationships. Forests doi: 10.3390/f11121250 www PDF
  10. Pompa-García, M., Hevia, A. and Camarero, J.J. 2020. Minimum and maximum wood density as proxies of water availability in two Mexican pine species coexisting in a seasonally dry area. Trees 35: 597-607. PDF
  11. Rubio-Cuadrado, A., Gómez, C., Rodríguez-Calcerrada, J., Perea, R., Gordaliza, G.G., Camarero, J.J., Montes, F. and Gil, L. 2020. Differential response of oak and beech to late frost damage: an integrated analysis from organ to forest. Agricultural and Forest Meteorology DOI:10.1016/j.agrformet.2020.108243. PDF
  12. Camarero, J.J. and Ortega, M.A. 2020. Tree growth in the aftermath of a flood: a tree-ring based reconstruction of the impacts of the 1996-Biescas catastrophe. Dendrochronologia 65: 125783. PDF
  13. Bader, M.Y., Llambí, L.-D., Case, B.S., Buckley, H.L., Toivonen, J.M., Camarero, J.J., Cairns, D.M., Brown, C.D., Wiegand,T. and Resler, L. M. 2020. A global framework for linking alpine-treeline ecotone patterns to underlying processes. Ecography (in press). PDF
  14. Lu, X., Liang, E., Wang, Y., Babst, F. and Camarero, J.J. 2020. Mountain treelines climb slowly despite rapid climate warming. Global Ecology and Biogeography DOI: 10.1111/geb.13214. PDF
  15. Batllori, E., Lloret, F., Aakala, T., Anderegg, W.R.L.,…Camarero, J.J., Colangelo, M….Zeeman, B. 2020. Forest and woodland replacement patterns following drought-related mortality. Proceedings of the National Academy of Sciences of the United States of America 10.1073/pnas.2002314117. PDF
  16. Sanmiguel-Vallelado, A., Camarero, J.J., Morán-Tejeda, E., Gazol, A., Colangelo, M., Alonso-González, E. and López-Moreno, J.I. 2020. Snow dynamics influence tree growth by controlling soil temperature in mountain pine forests. Agricultural and Forest Meteorology 296: 108205. PDF
  17. Camarero, J.J., Sánchez-Salguero, R., Sangüesa-Barreda, G., Lechuga, V., Viñegla, B., Seco, J.I., Taïqui, L., Carreira, J.A. and Linares, J. C. 2020. Drought, axe and goats. More variable and synchronized growth forecasts worsening dieback in Moroccan Atlas cedar forests. Science of the Total Environment 142752. PDF
  18. Rubio-Cuadrado, A., Camarero, J.J., Gordaliza, G.G, Cerioni, M., Montes, F. and Gil, L. 2020. Competition overrides climate as trigger of growth decline in a mixed Fagaceae Mediterranean rear-edge forest. Annals of Forest Science 77: 94. PDF
  19. Molina-Valero, J.A., Camarero, J.J., Álvarez-González, J.G., Cerioni, M., Hevia, A., Sánchez-Salguero, R., Martín-Benito, D. and Pérez-Cruzado, C. 2020. Mature forests hold maximum live biomass stocks. Forest Ecology and Management 480: 118635. PDF
  20. Serra-Maluquer, X., Gazol, A., Igual, J.M. and Camarero, J.J. 2020. Silver fir growth responses to drought depend on interactions between tree characteristics, soil and neighbourhood features. Forest Ecology and Management 480: 118625. PDF
  21. Navarro-Cerrillo, R.M., Gazol, A., Rodríguez-Vallejo, G., Manzanedo, R.D., Palacios-Rodríguez, G. and Camarero, J.J. 2020. Linkages between climate, radial growth and defoliation in Abies pinsapo forests from Southern Spain. Forests 11: 1002 www PDF
  22. Rodriguez-Vallejo, C., Navarro-Cerrillo, R.M., Manzanedo, R.D., Palacios Rodriguez, G., Gazol, A. and Camarero, J.J. 2020. High resilience, but low viability, of pine plantations in the face of a shift towards a drier climate. Forest Ecology and Management 479: 118537. PDF
  23. Camarero, J.J., Collado, E., Martínez-de-Aragón, J., De-Miguel, S., Büntgen, U., Martinez-Peña, F., Martín-Pinto, P., Ohenoja, E., Romppanen, T., Salo, K., Oria-de-Rueda, J.A. and Bonet, J.A. 2020. Associations between climate and earlywood and latewood width in boreal and Mediterranean Scots pine forests. Trees doi: 10.1007/s00468-020-02028-0. PDF
  24. Castaño, C., Camarero, J.J., Zas, R., Sampedro, L., Bonet, J.A., Alday, J.G. and Oliva, J. 2020. Insect defoliation is linked to a decrease in soil ectomycorrhizal biomass and shifts in needle endophyti communities. Tree Physiology 40: 1712-1725. PDF
  25. Rubio-Cuadrado, A., Camarero, J.J., Gómez, C., Cañellas, I., Aulló-Maestro, I., Gil, L. and Montes, F. 2020. Scots pine plantations growth adaptation to climate warming in locations at the southernmost distribution limit of the species. Dendrochronologia 63: 125745. PDF
  26. López-Sáez, J.A., Camarero, J.J., Schaad-Abel, D., Luelmo-Lautenschlaeger, R., Pérez-Díaz, S., Alba-Sánchez, F. and Carrión, J. 2020. Don’t lose sight of the forest for the trees! Discerning Iberian pine communities by means of pollen-vegetation relationships. Review of Palaeobotany and Palynology 281: 104285. PDF
  27. Garcia-Barreda, S. and Camarero, J.J. 2020. Tree ring and water deficit indices as indicators of drought impact on black truffle production in Spain. Forest Ecology and Management 475: 118438. PDF
  28. Camarero, J.J., Sánchez-Salguero, R., Ribas, M., Touchan, R., Andreu-Hayles, L., Dorado-Liñán, I., Meko, D.M. and Gutiérrez, E. 2020. Biogeographic, atmospheric and climatic factors influencing tree growth in Mediterranean Aleppo pine forests. Forests 11: 736. PDF
  29. Huang, J.-G., Ma, Q., Rossi, S., Biondi, F., Deslauriers, A., Fonti, P., Liang, E., Mäkinen, H., Oberhuber, W., Rathgeber, C.B.K., Tognetti, R., Treml, V., Yang, B., Antonucci, S., Bergeron, Y., Camarero, J.J., Campelo, F., Čufar, K., Cuny, H.E., De Luis, M., Giovannelli, A., Gričar, J., Gruber, A., Gryc, V., Güney, A., Guo, X., Huang, W., Ivanova, A., Jyske, T., Kašpar, K., King, G., Krause, C., Lombardi, F., Martinez, E., Morin, H., Nöjd, P., Peters, R., Prislan, P., Saracino, A., Swidrak, I., Vavrčík, H., Vieira, J., Yu, B., Zhai, L., Zhang, S., Zeng, Q. and Ziaco, E. 2020. Photoperiod and temperature as dominant environmental drivers triggering secondary growth resumption in Northern Hemisphere conifers. Proceedings of the National Academy of Sciences of the United States of America doi: 10.1073/pnas.2007058117. PDF
  30. Camarero, J.J., Pueyo, E.L., Sánchez-Moreno, E.L. and Calvín, P. 2020. Dating lightning: dendrochronological and magnetic analyses of lightning scars. Dendrochronologia 62: 125727. PDF
  31. Fu, T., Liang, E., Lu, X., Gao, S., Zhang, L., Zhu, H., Rossi, S. and Camarero, J.J. 2020. Tree growth responses and resilience after the 1950-Zayu-Medog earthquake, southeast Tibetan Plateau. Dendrochronologia 62: 125724. PDF
  32. Sánchez-Salguero, R., Colangelo, M., Matías, L., Ripullone, F. and Camarero, J.J. 2020. Shifts in growth responses to climate and surpassed drought-vulnerability thresholds characterize dieback in two Mediterranean deciduous oaks. Forests 11: 714. doi:10.3390/f11070714 www PDF
  33. Camarero, J.J., Gazol, A., Sánchez-Salguero, R., Sangüesa-Barreda, G., Díaz-Delgado, R. and Casals, R. 2020. Dieback and mortality of junipers caused by drought: dissimilar growth and wood isotope patterns preceding shrub death. Agricultural and Forest Meteorology 291: 108078. PDF
  34. Sangüesa-Barreda, G., Esper, E., Büntgen, U., Camarero, J.J., Di Filippo, A., Baliva, M. and Piovesan, G. 2020. Climate-human interactions contributed to historical forest recruitment dynamics in Mediterranean subalpine ecosystems. Global Change Biology doi: 10.1111/gcb.15246. PDF
  35. Lu, X., Huang, R., Wang, Y., Zhang, B., Zhu, H., Camarero, J.J., and Liang, E. 2020. Spring hydroclimate reconstruction on the south-central Tibetan Plateau inferred from Juniperus pingii var. wilsonii shrub rings since 1605. Geophysical Research Letters 10.1029/2020GL087707. PDF
  36. González-Díaz, P., Gazol, A., Valbuena-Carabaña, M., Sangüesa-Barreda, G., Moreno Urbano, A., Zavala, M.A. and Camarero, J.J. 2020. Remaking a stand: links between genetic diversity and tree growth in expanding Mountain pine populations. Forest Ecology and Management 472: 118244. PDF
  37. Gazol, A.,  Camarero, J.J., Sánchez-Salguero, R., Vicente-Serrano, S.M., Serra-Maluquer, X., Gutiérrez, E., De Luis, M., Sangüesa-Barreda,G., Novak, K., Rozas, V., Tíscar, P.A., Linares, J.C., Martínez del Castillo, E., Ribas, M., García-González, I., Silla, F., Camisón, A., Génova, M., Olano, J.M., Hereş, A.-M., Curiel Yuste, J., Longares, L.A., Hevia, A., Tomàs-Burguera, M. and Galván, J.D. 2020. Drought legacies are short, prevail in dry conifer forests and depend on growth variability. Journal of Ecology doi: 10.1111/1365-2745. PDF
  38. Bose, A.K., Gessler, A., Bolte, A., Bottero, A., Buras, A., Cailleret, M., Camarero, J.J., Haeni, M., Hereş, A.-M., Hevia, A., Lévesque, M., Linares, J.C., Martinez-Vilalta, J., Matías, L. Menzel, A., Sánchez-Salguero, R., Saurer, M., Vennetier, M., Ziche, D. and Rigling, A. 2020. Growth and resilience responses of Scots pine to extreme droughts across Europe depend on pre-drought growth conditions. Global Change Biology doi: 10.1111/gcb.15153. PDF
  39. Pandey, J., Sigdel, S.R., Lu, X., Salerno, F., Dawadi, B., Liang, E., and Camarero, J.J. 2020. Early growing-season precipitation drives radial growth of alpine juniper shrubs in the central Himalayas. Geografiska Annaler: Series A, Physical Geography DOI: 10.1080/04353676.2020.1761097. PDF
  40. Miranda, A., Lara, A., Altamirano, A., Di Bella, C.M., González, M.E. and Camarero, J.J. 2020. Forest browning trends in response to drought in a highly threatened Mediterranean landscape of South America. Ecological Indicators 115: 106401. PDF
  41. Manrique-Alba, A., Beguería. S., Molina, A.J., González-Sanchis, M., Tomàs-Burguera, M., del Campo, A.D., Colangelo, M. and Camarero, J.J. 2020. Long-term thinning effects on tree growth, drought response and water use efficiency at two Aleppo pine plantations in Spain. Science of the Total Environment 728: 138536. PDF
  42. Alday, J.G., Camarero, J.J., Revilla, J. and Resco de Dios, V. 2020. Similar diurnal, seasonal and annual rhythms in radial root expansion across two coexisting Mediterranean oak species. Tree Physiology 40: 956-968. PDF
  43. Camarero, J.J. and Hevia, A. 2020. Links between climate, drought and minimum wood density in conifers. IAWA J. 41: 236-255. PDF
  44. Kiorapostolou, N., Camarero, J.J., Carrer, M., Sterck, F., Brigita, B., Sangüesa-Barreda, G. and Petit, G. 2020. Scots pine trees react to drought by increasing xylem and phloem conductivities. Tree Physiology 40: 774–781. PDF
  45. Gao, S., Zhou, T., Yi, C., Shi, P., Fang, W., Liu, R., Liang, E. and Camarero, J.J. 2020. Asymmetric impacts of dryness and wetness on tree growth and forest coverage. Agricultural and Forest Meteorology 288-289: 107980 PDF
  46. Gazol, A., Sangüesa-Barreda, G. and Camarero, J.J. 2020. Forecasting forest vulnerability to drought in Pyrenean Silver fir forests showing dieback. Frontiers in Forests and Global Change doi: 10.3389/ffgc.2020.00036 www PDF
  47. García-Cervigón, A., Fajardo, A., Caetano-Sánchez, C., Camarero, J.J. and Olano, J.M. 2020. Xylem anatomy needs to change, so that conductivity can stay the same: xylem adjustments across elevation and latitude in Nothofagus pumilio. Annals of Botany 125: 1101–1112. PDF
  48. Ripullone, F., Camarero, J.J., Colangelo, M. and Voltas, J. 2020. Variation in the access to deep soil water pools explains tree-to-tree differences in drought-triggered dieback of Mediterranean oaks. Tree Physiology 40: 591–604. PDF
  49. Sánchez-Salguero, R. and Camarero, J.J. 2020. Greater sensitivity to hotter droughts underlies juniper dieback and mortality in Mediterranean shrublands. Science of the Total Environment (in press) PDF
  50. Sigdel, S.R., Liang, E., Wang, Y., Dawadi, B. and Camarero, J.J. 2020. Tree-to-tree interactions slow down Himalayan treeline shifts as inferred from tree spatial patterns. Journal of Biogeography 47:1816–1826. PDF
  51. Sanmiguel-Vallelado, A., López-Moreno, J.I., Morán-Tejeda, E., Alonso-González, E., Navarro-Serrano, F.M., Rico, I. and Camarero, J.J. 2020. Variable effects of forest canopies on snow processes in a valley of the central Spanish Pyrenees. Hydrological Processes 34: 2247-2262. doi 10.1002/hyp.13721. PDF
  52. García-Barreda, S., Camarero, J.J., Vicente-Serrano, S.-M. and Serrano-Notivoli, R. 2020. Variability and trends of black truffle production in Spain (1970-2017): linkages to climate, host growth, and human factors. Agricultural and Forest Meteorology 287: 107591 PDF
  53. Alfaro-Sánchez, R., Camarero, J.J., Querejeta, J.I., Sagra, J., Moya, D. and Rodríguez-Trejo, D.A. 2020. Volcanic activity signals in tree-rings at the treeline of the Popocatépetl, Mexico. Dendrochronologia doi 10.1016/j.dendro.2020.125663 PDF
  54. DeSoto, L., Cailleret, M., Sterck, F., Jansen, S., Kramer, K., Robert, E., Aakala, T., Amoroso, M., Bigler, C., Camarero, J.J., Cufar, K., Gea-Izquierdo, G., Gillner, S., Haavik, L., Heres, A.-M., Kane, J., Kharuk, V., Kitzberger, T., Klein, T., Levanic, T., Linares, J.C., Makinen, H., Oberhuber, W., Papadopoulos, A., Rohner, B., Sangüesa-Barreda, G., Stojanovic, D., Suarez, L.M., Villalba, R. and Martinez-Vilalta, J. 2020. Low growth resilience to drought is related to future mortality risk in trees. Nature Communications 11: 545. doi 10.1038/s41467-020-14300-5. www PDF


  1. Navarro-Cerrillo, R.N., Manzanedo, R.D., Rodríguez, C., Gazol, A., Palacios, G. and Camarero, J.J.
    2019. Competition modulates the response of growth to climate in pure and
    mixed Abies pinsapo subsp. maroccana forests in northern Morocco. Forest Ecology and Management 459 PDF
  2. Gazol, A., Hernández-Alonso, R. and Camarero, J.J. 2019. Patterns and drivers of pine processionary moth defoliation in Mediterranean mountain forests. Frontiers in Ecology and Evolution doi 10.3389/fevo.2019.00458 www PDF
  3. García-Cervigón, A.I., Camarero, J.J., Cueva, E., Espinosa, C.I. and Escudero, A. 2019. Climate seasonality and tree growth strategies in a tropical dry forest. Journal of Vegetation Science. DOI: 10.1111/jvs.12840 PDF
  4. Kattge, J., Bönisch, G., Díaz, S., Lavorel, S.,…Camarero, J.J.…and Wirth, C.
    2019. TRY plant trait database – enhanced coverage and open access. Global Change Biology 26: 119-188. PDF
  5. Lavergne, A., Voelker, S., Csank, A., Graven, H., de Boer, H., Daux, V., Robertson, I., Dorado-Liñan, I., Martínez-Sancho, E., Battipaglia, G., Bloomfield, K., Still, C., Meinzer, F., Dawson, T., Clisby, R., Camarero, J.J., Fang, Y., Menzel, A., Keen, R., Roden, J. and Prentice, C. 2019. Historical changes in the stomatal limitation of photosynthesis: empirical support for an optimality principle. New Phytologist 225: 2484–2497. PDF
  6. Ruiz-Benito, P., Vacchiano, G., Lines, E.R., Reyer, C.P.O., Ratcliffe, S., Morin, X., Hartig, F., Mäkelä, A., Yousefpour, R., Chaves, J.E., Palacios-Orueta, A., Benito-Garzón, M., Morales-Molino, C., Camarero, J.J., Jump, A.S., Kattge, J., Lehtonen, A., Ibrom, A., Owen, H.J.F. and Zavala, M.A. 2019. Available and missing data to model impact of climate change on European forests. Ecological Modelling 416 PDF
  7. Slette, I., Smith, M., Knapp, A., Vicente-Serrano, S.M., Camarero, J.J. and Beguería, S.
    2019. Standardized metrics are key for assessing drought severity. Global Change Biology DOI: 10.1111/gcb.14899 PDF
  8. Du, Q., Rossi, S., Lu, X., Wang, Y., Zhu, H., Liang, E. and Camarero, J.J. 2019. Negative growth responses to temperature of sympatric species converge under warming conditions in the southeastern Tibetan Plateau. Trees DOI: 10.1007/s00468-019-01924-4 PDF
  9. Vicente-Serrano, S.M., Martín-Hernández, N., Camarero, J.J., Gazol, A., Sánchez-Salguero, R.,  Peña-Gallardo, M., El Kenawy, A., Domínguez-Castro, F., Tomas-Burguera, M., Gutiérrez, E., De Luis, M., Sangüesa-Barreda, G., Novak, K., Rozas, V., Tíscar, P.A., Linares, J.C., Martínez del Castillo, E., Ribas, M., García-González, I., Silla, F.,  Camisón, A., Génova, M., Olano, J.M., Longares, L.A., Hevia, A. and Galván, J.D. 2019. Linking tree-ring growth
    and satellite-derived gross primary growth in multiple forest biomes.
    Temporal-scale matters. Ecological Indicators 108 PDF
  10. Rita, A., Camarero, J.J., Nolè, A., Borghetti, M., Brunetti, M., Pergola, N., Serio, C., Vicente-Serrano, S.M., Tramutoli, V. and Ripullone, F.
    2019. The impact of drought spells on forests depends on site conditions: the case of 2017 summer heat wave in southern Europe. Global Change Biology DOI: 10.1111/gcb.14825 PDF
  11. Sangüesa-Barreda, G., Camarero, J.J., Sánchez-Salguero, R., Gutiérrez, E., Linares, J.C., Génova, M., Ribas, M., Tíscar, P.A. and López-Sáez, J.A. 2019. Droughts and climate warming desynchronize Black pine growth across the Mediterranean Basin. Science of the Total Environment 697 PDF
  12. Pacheco, A., Camarero, J.J., Pompa-García, M., Battipaglia, G., Voltas, J. and Carrer, M. 2019. Growth, wood anatomy and stable isotopes shows species-specific couplings in three Mexican conifers inhabiting drought-prone areas. Science of the Total Environment 698 PDF
  13. Gut, U., Árvai, M., Bijak, S., Camarero, J.J., Cedro, A., Cruz-García, R., Garamszegi, B., Hacket-Pain, A., Hevia, A., Huang, W., Isaac-Renton, M., Kaczka, R., Kazimirović, M., Kędziora, W., Kern, Z., Klisz, M., Kolar, T., Körner, M., Kuznetsova, V., Montwé, D., Petritan, A.M., Petritan, I.C., Plavcová, L., Rehschuh, R., Rocha, E., Rybníček, M., Sánchez-Salguero, R., Schröder, J., Schwab, N., Stajic, B., Tomusiak, R., Wilmking, M., Sass-Klaassen, U., and Buras, A. 2019. No systematic effects of sampling direction on climate-growth relationships in a large-scale, multi-species tree-ring data set. Dendrochronologia 57 PDF
  14. Gazol, A., Camarero, J.J., Colangelo, M., De Luis, M., Martínez del Castillo, E. and Serra-Maluquer, X. 2019. Summer drought and spring frost, but not their interaction, constrain European beech and Silver fir growth in their southern distribution limits. Agricultural and Forest Meteorology 278 PDF
  15. Sanmiguel-Vallelado, A., Camarero, J.J., Gazol, A., Morán-Tejeda, E., Sangüesa-Barreda, G., Alonso-González, E., Gutiérrez, E., Alla, A.Q., Galván, J.D., and López-Moreno, J.I. 2019. Detecting snow-related signals in radial growth of Pinus uncinata mountain forests. Dendrochronologia 57 PDF
  16. Hevia, A., Sánchez-Salguero, R., Camarero, J.J., Querejeta, J.I., Sangüesa-Barreda, G. and Gazol, A. 2019. Long-term nutrient imbalances linked to drought-triggered forest dieback. Science of the Total Environment 690: 1254-1267. PDF
  17. Collado, E.A., Bonet, J.A., Camarero, J.J., Egli, S., Peter, M., Salo, K.E., Martínez-Peña, F., Ohenoja, E., Martín-Pinto, P., Primicia, I., Büntgen, U., Kurttila, M., Oria-de-Rueda, J.A., Martínez-de-Aragón, J., Miina, J. and De-Miguel, S. 2019. Mushroom productivity trends in relation to tree growth and climate across different European forest biomes. Science of the Total Environment 689: 602-615. PDF
  18. González-Casares, M., Camarero, J.J., Colangelo, M., Rita, A. and Pompa-García, M. 2019. High responsiveness of wood anatomy to water availability and drought near the equatorial rear-edge of Douglas fir. Canadian Journal of Forest Research 49: 1114-1123. PDF
  19. Camarero, J.J., González de Andrés, E., Sangüesa-Barreda, G., Rita, A. and Colangelo, M. 2019. Long- and short-term impacts of a defoliating moth plus mistletoe on tree growth, wood anatomy and water-use efficiency. Dendrochronologia 56 PDF
  20. Büntgen, U., Krusic, P.J., Piermattei, A., Coomes, D.A., Esper, J., Myglan, V.S., Kirdyanov, A.V., Camarero, J.J., Crivellaro, A. and Körner, Ch. 2019. Limited capacity of tree growth to mitigate the global greenhouse effect under predicted warming.
    Nature Communications 10: 2171. www PDF
  21. Sidor, C.G., Camarero, J.J., Popa, I., Badea, O., Apostol, E.N. and Vlad, R. 2019. Forest vulnerability to extreme climatic events in Romanian Scots pine forests. Science of the Total Environment 678: 721-727. PDF
  22. Navarro-Cerrillo, R.M., Sarmoum, M., Gazol, A., Abdoun F. and Camarero, J.J. 2019. The decline of Algerian Cedrus atlantica forests is driven by a climate shift towards drier conditions. Dendrochronologia 55: 60-70. PDF
  23. Carnicer, J., Domingo-Marimon, C., Ninyerola, M., Camarero, J.J., Bastos, A., López-Parages, J., Rodríguez-Fonseca, B., Lenton, T.M., Dakos, V., Ribas, M., Gutiérrez, E., Peñuelas, J. and Pons, X.
    2019. Regime shifts of Mediterranean forest carbon
    uptake and reduced resilience driven by multidecadal ocean surface temperature. Global Change Biology DOI: 10.1111/gcb.14664. PDF
  24. Wang, Y., Sylvester, S.P., Lu, X., Dawadi, B., Sigdel, S.R., Liang, E. and Camarero, J.J. 2019. The stability of spruce treelines on the eastern Tibetan Plateau over the last century is explained by pastoral disturbance. Forest Ecology and Management 442: 34-45. PDF
  25. De Micco, V., Carrer, M., Rathgeber, C.B.K., Camarero, J.J., Voltas, J., Cherubini, P., and Battipaglia, G. 2019. From xylogenesis to tree-rings: wood traits to investigate tree-response to environmental changes. IAWA Journal 40: 155-182. PDF
  26. Fajardo, A., Gazol, A., Mayr, C. and Camarero, J.J. 2019. Recent decadal drought reverts warming–triggered growth enhancement in contrasting climates in the southern Andes treeline. Journal of Biogeography 46: 1367-1379. PDF
  27. Wang, Y., Case, B., Lu, X., Ellison, A.M., Peñuelas, J., Zhu, H., Liang, E. and Camarero, J.J. 2019. Fire facilitates warming-induced upward shifts of alpine treelines by altering interspecific interactions. Trees DOI: 10.1007/s00468-019-01841-6 PDF
  28. Camarero, J.J., Sangüesa-Barreda, G., Pérez-Díaz, S., Montiel-Molina, C., Seijo, F. and López-Sáez, J.A. 2019. Abrupt regime shifts in post-fire resilience of Mediterranean mountain pinewoods are fueled by land use. International Journal of Wildland Fire, doi 10.1071/WF18160. PDF
  29. Lu, X., Liang, E., Wang, Y., Babst, F., Leavitt, S.W. and Camarero, J.J. 2019. Past the climate optimum: Recruitment is declining at the world’s highest juniper shrublines on the Tibetan Plateau. Bulletin of the Ecological Society of America 100: e01497. PDF
  30. Vitali, A., Garbarino, M., Camarero, J.J., Malandra, F., Toromani, E., Spalevic, V., Curovic, M. and Urbinati, C. 2019. Pine recolonization dynamics in Mediterranean human-disturbed treeline ecotones. Forest Ecology and Management 435: 28-37. PDF
  31. Cailleret, M., Dakos, V., Jansen, S., Robert, E.M., Aakala, T., Amoroso, M.M., Antos, J.A., Bigler, C., Bugmann, H., Caccianiga, M., Camarero, J.J. et al. 2019. Early-warning signals of individual tree mortality based on annual radial growth. Frontiers in Plant Science doi 10.3389/fpls.2018.01964. www PDF
  32. Serra-Maluquer, X., Gazol, A., Sangüesa-Barreda, G., Sánchez-Salguero, R., Rozas, V., Colangelo, M., Gutiérrez, E. and Camarero, J.J. 2019. Geographically structured growth decline of rear-edge Iberian Fagus sylvatica forests after the 1980s shift toward a warmer climate. Ecosystems doi 10.1007/s10021-019-00339-z PDF
  33. Delpierre, N., Lireux, S., Hartig, F., Camarero, J.J. et al. 2019. Chilling and forcing temperatures interact to predict the onset of wood formation in Northern Hemisphere conifers. Global Change Biology doi 10.1111/gcb.14539. PDF


  1. Brodde, L., Adamson, K., Camarero, J.J., Castaño, C., Drenkhan, R., Lehtijarvi, A., Luchi, N., Migliorini, D., Sánchez-Miranda, A., Stenlid, J., Özdağ, S. and Oliva, J. 2018. Diplodia tip blight on its way to the north: drivers of disease emergence in Northern Europe. Frontiers in Plant Science doi: 10.3389/fpls.2018.01818. www PDF
  2. Camarero, J.J. and Ortega, M.A. 2018. Sancho, the oldest known Iberian shrub. Dendrochronologia 53: 32-36. PDF
  3. Navarro, R.M., Sánchez-Salguero, R., Rodríguez, C., Duque, J., Moreno-Rojas, J.M., Palacios-Rodríguez, G. and Camarero, J.J. 2018. Is thinning an alternative when trees could die in response to drought?
    The case of planted Pinus nigra and P. sylvestris stands in southern
    Spain. Forest Ecology and Management 433: 313-324. PDF
  4. Sánchez-Salguero, R., Camarero, J.J., Gutiérrez, E., Gazol, A., Sangüesa-Barreda, G., Moiseev, P. and Linares, J.C. 2018. Climate warming alters age-dependent growth sensitivity to temperature in Eurasian alpine treelines. Forests 9: 688 PDF www
  5. Barbeta, A., Camarero, J.J., Sangüesa-Barreda, G., Muffler, L. and Peñuelas, J. 2018. Contrasting effects of fog frequency on the radial growth of two tree species in a Mediterranean-temperate ecotone. Agricultural and Forest Meteorology 264: 297-308. PDF
  6. Colangelo, M., Camarero, J.J., Borghetti, M., Gentilesca, T., Oliva, J., Ripullone, F., and Redondo, M.A. 2018. Drought and Phytophthora are associated with the decline of oak species in southern Italy. Frontiers in Plant Science doi 10.3389/fpls.2018.01595 PDF www
  7. Lu, X., Liang. E., Wang, Y., Babst, F., Leavitt, S, and Camarero, J.J. 2018. Past the climate optimum: Recruitment is declining at the world’s highest juniper shrublines on the Tibetan Plateau. Ecology doi: 10.1002/ecy.2557. PDF
  8. Camarero, J.J., Álvarez-Taboada, F., Hevia, A., and Castedo-Dorado, F. 2018. Radial growth and wood density reflect the impacts and susceptibility to defoliation by gypsy moth and climate in radiata pine. Frontiers in Plant Science PDF www
  9. Gazol, A., Camarero, J.J., Sangüesa-Barreda, G., and Vicente-Serrano, S.M. 2018. Post-drought resilience after forest die-off: shifts in regeneration, composition, growth and productivity. Frontiers in Plant Science PDF www
  10. Camarero, J.J., Sánchez-Salguero, R., Sangüesa-Barreda, G. and Matías, L. 2018. Tree species from contrasting hydrological niches
    show divergent growth and water-use efficiency. Dendrochronologia 52: 87-95. PDF
  11. Garcés-Pastor, S., Gutiérrez, E., Martínez-Sancho, E., Dorado-Liñán, I., Camarero, J.J., Cañellas-Boltà, N. and Vegas-Vilarrúbia, T. 2018. Subalpine forest dynamics reconstructed throughout the last 700 years in the Central Pyrenees by means of tree rings and pollen. The Holocene PDF
  12. Acosta-Hernández, A.C., Camarero, J.J. and Pompa-García, M. 2018. Seasonal growth responses to climate in wet and dry conifer forests. IAWA Journal 40: 311-330. PDF
  13. Camarero, J.J. 2018. Linking functional traits and climate-growth
    relationships in Mediterranean species through wood density. IAWA Journal  40: 215-240. PDF
  14. Hernández, L., Camarero, J.J., Gil-Pelegrín, E., Saz, M.A., Cañellas, I. and Montes, F. 2018. Biotic interactions and increasing aridity shape the altitudinal shifts of marginal Pyrenean silver fir populations in Europe. Forest Ecology and Management 432: 558-567. PDF
  15. Sánchez-Salguero, R., Camarero, J.J., Hevia, A., Sangüesa-Barreda, G., and Gutiérrez, E. 2019. Testing annual tree-ring chemistry by X-ray
    fluorescence for dendroclimatic
    studies in high-elevation forests from the Spanish Pyrenees. Quaternary International 514: 130-140. PDF
  16. Espinosa, C.I., Camarero, J.J., and Gusmán, A.A. 2018. Site-dependent growth responses to climate in two major tree species from tropical dry forests of southwest Ecuador. Dendrochronologia 52: 11-19. PDF
  17. Peña-Gallardo, M., Vicente-Serrano, S.M., Camarero, J.J., Gazol, A., Sánchez-Salguero, R., Domínguez-Castro, F., El Kenawy, A., Beguería-Portugés, S., Gutiérrez, E., de Luis, M., Sangüesa-Barreda, G., Novak, K., Rozas, V., Tíscar, P.A., Linares, J.C., Martínez del Castillo, E., Ribas, M., García-González, I., Silla, F., Camisón, A., Génova, M., Olano, J.M., Longares, L.A., Hevia, A. and Galván, J.D. 2018. Drought sensitiveness on NDVI and forest growth in peninsular Spain
    and the Balearic Islands. Forests 9: 524 PDF www
  18. Sidgel, S.R., Wang, Y., Camarero, J.J., Zhu, H., Liang, E. and Peñuelas, J. 2018. Moisture-mediated responsiveness of treeline shifts to global warming in the Himalayas. Global Change Biology DOI: 10.1111/gcb.14428. PDF
  19. Fernández-Manjarrés, J., Ruiz-Benito, P., Zavala, M.A., Camarero, J.J., Proença, V., Navarro, L., Sansilvestri, R., Marqués, L., Temunovič, M., Bertelsmeier, C., Pulido, F., Drobinski, J.P., Roturier, S., Benito-Garzón, M., García de Cortazar-Atauri, I., Simon, L., Dupas, S., Levrel, H., and Sautier, M. 2018. Forest adaptation to climate change along steep ecological gradients:
    The case of the Mediterranean-temperate transition in South-Western Europe. Sustainability 10: 3065 PDF www
  20. Rubio-Cuadrado, A., Camarero, J.J., Aspizua, R., Sánchez-González, M., Gil, L. and Montes, F. 2018. Abiotic factors modulate post-drought growth resilience of
    Scots pine plantations and rear-edge Scots pine and oak forests. Dendrochronologia 51: 54-65. PDF
  21. García-Cervigón, A., Linares, J.C., García-Hidalgo, M., Camarero, J.J. and
    Olano, J.M. 2018. Growth delay by winter precipitation could hinder Juniperus sabina persistence under increasing summer drought. Dendrochronologia 51: 22-31. PDF
  22. Herrera-Soto, G., González-Cásares, M., Pompa-García, M., Camarero, J.J. and Solís-Moreno, R. 2018. Response of Pinus cembroides growth to hydroclimatic variability at the distribution limits of the species. Forests 9: 440; doi:10.3390/f9070440 PDF www
  23. Olson, M.E., Soriano, D., Rosell, J.A., Anfodillo, T., Donoghue, M.J., Edwards, E.J., León-Gómez, C., Dawson, T., Camarero, J.J., Castorena, M., Echeverría, A., Espinosa, C.I., Fajardo, A., Gazol, A., Isnard, S., Lima, R.S., Marcati, C.R. and Méndez-Alonzo, R. 2018. Plant height and hydraulic vulnerability to drought and cold. Proceedings of the National Academy of Sciences of the United States of America doi:10.1073/pnas.1721728115 PDF
  24. Rubio-Cuadrado, A., Camarero, J.J., del Río, M.,
    Sánchez-González, M., Ruiz-Peinado, R., Bravo-Oviedo, A., Gil, L. and
    Montes, F. 2018. Drought modifies tree competitiveness in an oak-beech temperate forest. Forest Ecology and Management 429: 7-17. PDF
  25. Sigdel, S.R., Dawadi, B., Camarero, J.J., Liang, E. and Leavitt, S.W. 2018. Moisture-limited tree growth for a subtropical Himalayan conifer forest in western Nepal. Forests 9: 340 PDF www
  26. Sangüesa-Barreda, G., Camarero, J.J., Esper, J., Galván. D. and Büntgen, U. 2018. A millennium-long perspective on high-elevation pine recruitment in the Spanish central Pyrenees. Canadian Journal of Forest Research (in press)
  27. Navarro-Cerrillo, R.M., Rodríguez-Vallejo, C., Silveiro, E., Hortal, A., Palacios-Rodríguez, G., Duque-Lazo, J. and Camarero, J.J. 2018. Cumulative drought stress leads to a loss of growth resilience and explains higher mortality in planted than in naturally regenerated Pinus pinaster stands. Forests 9: 358 PDF www
  28. Camarero, J.J., Gazol, A., Sangüesa-Barreda, G. and Fajardo, A. 2018. Coupled climate–forest growth shifts in the Chilean Patagonia are decoupled from trends in water–use efficiency. Agricultural and Forest Meteorology 259: 222-231. PDF
  29. Liu, B., Liang, E., Liu, K. and Camarero, J.J. 2018. Species- and elevation-dependent growth responses to climate warming of mountain forests in the Qinling Mountains, central China. Forests 9: 248 PDF www
  30. Rubio-Cuadrado, A., Camarero, J.J., del Río, M., Sánchez-González, M., Ruiz-Peinado, R., Bravo-Oviedo, A., Gil, L. and Montes, F. 2018. Long-term impacts of drought on growth and forest dynamics in a temperate beech-oak-birch forest. Agricultural and Forest Meteorology 259: 48-59. PDF
  31. Sangüesa-Barreda, G., Camarero, J.J., Pironon, S., Gazol, A., Peguero-Pina, J.J. and Gil-Pelegrín, E. 2018. Delineating limits: Confronting predicted climatic suitability to field performance in mistletoe populations. Journal of Ecology DOI: 10.1111/1365-2745.12968 PDF
  32. Gazol, A., Camarero, J.J., Jiménez, J.J., Moret-Fernández, D., López, M.V., Sangüesa-Barreda, G. and Igual, J.M. 2018. Beneath the canopy: linking drought-induced forest die off and changes in soil properties. Forest Ecology and Management 422: 294-302. PDF
  33. Collado, E., Camarero, J.J., Martínez de Aragón, J., Pemán, J., Bonet, J.A. and de-Miguel, S. 2018. Linking fungal dynamics, tree growth and forest management in a Mediterranean pine ecosystem. Forest Ecology and Management 422: 223-232. PDF
  34. Colangelo, M., Camarero, J.J., Ripullone, F., Gazol, A., Sánchez-Salguero, R., Oliva, J. and Redondo, M.A. 2018. Drought decreases growth and increases mortality of coexisting native and introduced tree species in a temperate floodplain forest. Forests 9: 205 PDF www
  35. Campelo, F., Gutiérrez, E., Ribas, M., Sánchez-Salguero, R., Nabais, C., and Camarero, J.J. 2018. The facultative bimodal growth pattern in Quercus ilex – a simple model to predict sub-seasonal and inter-annual growth. Dendrochronologia 49: 77-88. PDF
  36. Gazol, A., Camarero, J.J., Vicente-Serrano, S.M., Sánchez-Salguero, R., Gutiérrez, E., De Luis, M., Sangüesa-Barreda, G. et al. 2018. Forest resilience to drought varies across biomes. Global Change Biology DOI: 10.1111/gcb.14082 PDF
  37. Granda, E., Gazol, A. and Camarero, J.J. 2018. Functional diversity differently shapes growth resilience to drought for coexisting pine species. Journal of Vegetation Science DOI: 10.1111/jvs.12617 PDF
  38. Pacheco, A., Camarero, J.J. and Carrer M.  2018. Shifts of irrigation in Aleppo pine forests under semi-arid conditions reveal uncoupled growth and carbon storage and legacy effects on wood anatomy. Agricultural and Forest Meteorology 253-254: 225-232. PDF
  39. Palacio, S., Camarero, J.J., Maestro, M., Alla, A.Q., Lahoz, E., and Montserrat-Martí, G. 2018. Are storage and tree growth related? Seasonal nutrient and carbohydrate dynamics in evergreen and deciduous Mediterranean oaks. Trees 32: 777-790. PDF
  40. Caminero, L., Sánchez-Salguero, R., Génova, M., and Camarero, J.J. 2018. Growth responses to climate and drought at the southernmost European limit of Mediterranean Pinus pinaster forests. Dendrochronologia 48: 20-29. PDF
  41. Sánchez-Salguero, R., Camarero, J.J., Rozas, V., Génova, M., Olano, J.M., Arzac, A., Gazol, A., Caminero, L., Tejedor, E., de Luis M., and Linares, J.C. 2018. Resist, recover or both? Growth plasticity in response to drought is geographically structured and linked to intra-specific variability in Pinus pinaster. Journal of Biogeography DOI: 10.1111/jbi.13202 PDF
  42. Camarero, J.J., Sangüesa-Barreda, G., Montiel-Molina, C., Seijo, F., and López-Sáez, J.A. 2018. Past growth suppressions as proxies of fire incidence in relict Mediterranean black pine forests. Forest Ecology and Management 413: 9-20. PDF
  43. Trnka, M., Hayes, M., Jurečka, F., Bartošová, L., Anderson, M., Brázdil, R., Brown, J., Camarero, J.J., Cudlín, P., Dobrovolný, P., Eitzinger, J., Feng, S., Finnessey, T., Gregoric, G., Havlik, P., Hain, C., Holman, I., Johnson, D., Kersebaum, K., Ljungqvist, F., Luterbacher, J., Micale, F., Hartl-Meier, C., Možný, M., Nejedlik , P., Olesen, J., Ruiz-Ramos, M., Rötter, R., Senay, G., Vicente-Serrano, S.M., Svoboda, M., Susnika, A., Tadesse, T., Vizina, A., Wardlow, B., Büntgen, U., and Žalud, Z. 2018. Priority questions in multidisciplinary drought research. Climate Research
  44. Laskurain, N.A., Aldezabal, A., Odriozola, I., Camarero, J.J., and Olano, J.M. 2018. The role of tree context: variation in the individual growth response to climate among neighbourhoods and species in a secondary forest. Forests 9: 43; doi:10.3390/f9010043 PDF www
  45. Camarero, J.J., Gazol, A., Sangüesa-Barreda, G., Cantero, A., Sánchez-Salguero, R., Sánchez-Miranda, A., Granda, E., Serra-Maluquer, X. and Ibañez, R. 2018. Forest growth responses to drought at short- and long-term scales in Spain: squeezing the stress memory from tree rings. Frontiers in Ecology and Evolution 6: 9. doi: 10.3389/fevo.2018.00009 PDF www
  46. Cardil, A., Imbert, J.B., Camarero, J.J., Primicia, I. and Castillo, F.J.  2018. Temporal interactions among throughfall, type of canopy and thinning drive radial growth in an Iberian mixed pine-beech forest. Agricultural and Forest Meteorology 252: 62-74. PDF
  47. Ren, P., Rossi, S., Camarero, J.J., Ellison, A.M., Liang, E. and Peñuelas, J. 2018. Critical temperature and precipitation thresholds for the onset of xylogenesis of Juniperus przewalskii in a semi-arid area of the north-eastern Tibetan Plateau. Annals of Botany 121: 617-624. PDF
  48. Hevia, A., Sánchez-Salguero, R., Camarero, J.J., Buras, A., Sangüesa-Barreda, G., Galván, J.D., and Gutiérrez, E. 2018. Towards a better understanding of long-term wood-chemistry variations in old-growth forests: A case study on ancient Pinus uncinata trees from the Pyrenees. Science of the Total Environment 625: 220-232. PDF


  1. Candel-Pérez, D., Lo Y.-H., Blanco, J.A., Chiu, C.-M., Camarero, J.J., González de Andrés, E., Imbert, J.B. and Castillo, F.J. 2017. Drought-induced changes in wood density are not prevented by thinning. Forests 9: 4; doi: 10.3390/f9010004 PDF www
  2. Liang, E. and Camarero, J.J. 2017. Threshold-dependent and non-linear associations between temperature and tree growth at and below the alpine treeline. Trees 32: 661-662. PDF
  3. García-Barreda, S., Forcadell, R. Sánchez, S., Martín-Santafé, M., Marco, P., Camarero, J.J. and Reyna, S. 2017. Black truffle harvesting in Spanish forests: trends, current policies and practices, and implications on its sustainability. Environmental Management 61:535-544.  PDF 
  4. Granda, E., Alla, A.Q., Laskurain, N.A., Loidi, J., Sánchez-Lorenzo, A. and Camarero, J.J. 2017. Coexisting oak species, including rear-edge populations, buffer climate stress through xylem adjustments. Tree Physiology 38:159-172. PDF 
  5. Sánchez-Salguero, R., Camarero, J.J., Carrer, M., Gutiérrez, E., Alla, A.Q., Andreu-Hayles, L., Hevia, A., Koutavas, A., Martínez-Sancho, E., Nola, P., Papadopoulos, A., Pasho, E., Toromani, E., Carreira, J.A. and Linares, J.C. 2017. Climate extremes and predicted warming threaten Mediterranean Holocene firs forests refugia. Proceedings of the National Academy of Sciences of the United States of America doi:10.1073/pnas.1708109114PDF 
  6. Gazol, A., Moiseev, P. and Camarero, J.J. 2017. Changes in plant taxonomic and functional diversity patterns following treeline advances in the South Urals. Plant Ecology and Diversity 10: 283-292. PDF
  7. Vitali, A., Camarero, J.J., Garbarino, M., Piermattei, A. and Urbinati, C. 2017. Deconstructing human-shaped treelines: Microsite topography and distance to seed source shape Pinus nigra colonization of treeles areas in the Italian Apennines. Forest Ecology and Management 406: 37-45. PDF 
  8. Pompa-García, M., Camarero, J.J., Rodríguez-Tejo, D.A. and Vega-Nieva, D.J. 2017. Drought and spatiotemporal variability of forest fires across Mexico. Chinese Geographical Science 27 doi: 10.1007/s11769-017-0928-0. PDF
  9. Gentilesca, T., Camarero, J.J., Colangelo, M., Nolè, A. and Ripullone, F. 2017. Drought-induced oak decline in the western Mediterranean region: an overview on current evidences, mechanisms and management options to improve forest resilience. iForests 10: 796-806. PDF www
  10. Marqués, L., Madrigal-González, J., Zavala, M.A., Camarero, J.J. and Hartig, F. 2017. Last-century forest productivity in a managed dry-edge Scots pine population: the two sides of climate warming. Ecological Applications. PDF
  11. Alfaro-Sánchez, R., Camarero, J.J., Sánchez-Salguero, R., Trouet, V. and De Las Heras, J. 2017. How do droughts and wildfires alter seasonal radial growth in Mediterranean Aleppo pine forests? Tree-Ring Research 74: 1-14. PDF
  12. Pacheco, A., Camarero, J.J., Ribas, M., Gazol, A., Gutiérrez, E. and Carrer, M. 2017. Disentangling the climate-driven bimodal growth pattern in coastal and continental Mediterranean pine stands. Science of the Total Environment. PDF
  13. Pompa-García, M., Sánchez-Salguero, R. and Camarero, J.J. 2017. Observed and projected impacts of climate on radial growth of three endangered conifers in northern Mexico indicate high vulnerability of drought-sensitive species from mesic habitatsDendrochronologia 45: 145-155. PDF Supp Info
  14. Camarero, J.J. and Gutiérrez, E. 2017. Wood density of silver fir reflects drought and cold stress across climatic and biogeographic gradients. Dendrochronologia 45: 101-112. PDF
  15. Granda, E. and Camarero, J.J. 2017. Drought reduces growth and stimulates sugar accumulation: new evidence of environmentally driven non-structural carbohydrate use. Tree Physiology DOI: PDF
  16. Gazol. A., Sangüesa-Barreda, G., Granda, E. and Camarero, J.J. 2017. Tracking the impact of drought on functionally different woody plants in a Mediterranean scrubland ecosystem. Plant Ecology 218: 1009-1020. PDF
  17. González de Andrés, E., Camarero, J.J., Blanco, J.A., Imbert, J.B., Lo, Y.-H., Sangüesa-Barreda, G. and Castillo, F.J. 2017. Tree-to-tree competition in mixed European beech–Scots pine forests has different impacts on growth and water-use efficiency depending on site conditions. Journal of Ecology DOI 10.1111/1365-2745.12813. PDF
  18. Granda, E., Camarero, J.J. , Galván, J.D., Sangüesa-Barreda, G., Alla, A.Q., Gutiérrez, E., Dorado-Liñán, I.,  Andreu-Hayles, L., Labuhn, I., Grudd, H. and Voltas, J. 2017. Aged but withstanding: Maintenance of growth rates in old pines is not related to enhanced water-use efficiencyAgricultural and Forest Meteorology 243: 43-54. PDF
  19. Pompa-García, M., González-Cásares, M., Acosta-Hernández, A.C., Camarero, J.J. and Rodríguez-Catón, M. 2017. Drought influence over radial growth of Mexican conifers inhabiting mesic and xeric sites. Forests 8: 175; doi:10.3390/f8050175 PDF www
  20. Alfaro-Sánchez, R., Muller-Landau, H.C., Wright S.J. and Camarero, J.J. 2017. Growth and reproduction respond differently to climate in three Neotropical tree species. Oecologia DOI 10.1007/s00442-017-3879-3 PDF
  21. Cabral-Alemán, C., Pompa-García, M., Acosta-Hernández, A.C., Zúñiga-Vásquez, J.M. and Camarero, J.J. 2017. Earlywood and latewood widths of Picea chihuahuana show contrasting sensitivity to seasonal climate. Forests 8: 173, doi:10.3390/f8050160 PDF www
  22. Acosta-Hernández, A.C., Pompa-García, M. and Camarero, J.J. 2017. An updated review of dendrochronological investigations in Mexico, a megadiverse country with a high potential for tree-ring sciences. Forests 8: 160, doi:10.3390/f8050160 PDF www
  23. Camarero, J.J., Fernández-Pérez, L., Kirdyanov, A.V., Shestakova, T.A., Knorre, A.A., Kukarskih, V.V. and Voltas, J. 2017. Minimum wood density of conifers portrays changes in early season precipitation at dry and cold Eurasian regions. Trees doi 10.1007/s00468-017-1559-x. PDF
  24. Wang, Y., Liang, E., Sigdel, S.R., Liu, B. and Camarero, J.J. 2017. The coupling of treeline elevation and temperature is mediated by non-thermal factors on the Tibetan Plateau. Forests 8: 109, doi:10.3390/f8040109 PDF www
  25. Büntgen, U., Krusic, P.J., Verstege, A., Sangüesa-Barreda, G., Wagner, S., Camarero, J.J., Ljungqvist, F.C., Zorita, E., Oppenheimer, C., Konter, O., Tegel, W., Gärtner, H., Cherubini, P., Reinig, F. and Esper, J. 2017. New tree-ring evidence from the Pyrenees reveals Western Mediterranean climate variability since medieval times. Journal of Climate PDF
  26. Camarero, J.J. and Fajardo, A. 2017. Poor acclimation to current drier climate of the long-lived tree species Fitzroya cupressoides in the temperate rainforest of southern Chile. Agricultural and Forest Meteorology 239: 141-150. PDF
  27. Shestakova, T.A., Camarero, J.J., Ferrio, J.P., Knorre, A.A., Gutiérrez, E. and Voltas, J. 2017. Increasing drought effects on five European pines modulate Δ13C-growth coupling along a Mediterranean altitudinal gradient. Functional Ecology doi:10.1111/1365-2435.12857. PDF
  28. Touchan, R., Anchukaitis, K.J., Meko, D.M., Kerchouche, D., Slimani, S., Ilmen, R., Hasnaoui, F., Guibal, F., Camarero, J.J. et al. 2017. Climate controls on tree growth in the Western Mediterranean. The Holocene doi 10.1177/0959683617693901. PDF
  29. García-Cervigón, A.I., Camarero, J.J. and Espinosa, C.I. 2017. Intra-annual stem increment patterns and climatic responses in five tree species from an Ecuadorian tropical dry forest. Trees 31: 1057-1067. PDF
  30. Colangelo, M., Camarero, J.J., Battipaglia, G., Borghetti, M., De Micco, V., Gentilesca, T. and Ripullone, F. 2017. A multi-proxy assessment of dieback causes in a Mediterranean oak species. Tree Physiology 37: 617-631. PDF
  31. Colangelo, M., Camarero, J.J., Borghetti, M., Gazol, A. and Ripullone, F. 2017. Size matters a lot: drought-affected Italian oaks are smaller and show lower growth prior to tree death. Frontiers in Plant Science 8:135. doi: 10.3389/fpls.2017.00135 PDF www
  32. Sánchez-Salguero, R., Hevia, A., Camarero, J.J., Treydte, K., Frank, D., et al. 2017. An intensive tree-ring experience: Connecting education and research during the 25th European Dendroecological Fieldweek (Asturias, Spain). Dendrochronologia 42: 8093PDF
  33. Sánchez-Salguero, R., Camarero, J.J., Grau, J.M., De la Cruz, A., Gil, P.M., Minaya, M. and Fernández-Cancio, A. 2017. Analysing atmospheric processes and climatic drivers of tree defoliation to determine forest vulnerability to climate warming. Forests 8: 13, doi 10.3390/f8010013 PDF www
  34. Seijo, F., Millington, J.S.A., Gray, R., Hernández-Mateo, L., Sangüesa-Barreda, G. and Camarero, J.J. 2017. Divergent fire regimes in two contrasting Mediterranean chestnut forest landscapes. Human Ecology  doi 10.1007/s10745-016-9879-9 PDF
  35. Choury, Z., Shestakova, T.A., Himrane, H., Touchan, R., Kerchouche D., Camarero, J.J. and Voltas, J. 2017. Quarantining the Sahara desert: growth and water-use efficiency of Aleppo pine in the Algerian Green Barrier. European Journal of Forest Research 136: 139-152. PDF
  36. Gazol, A., Ribas, M., Gutiérrez, E. and Camarero, J.J. 2017. Aleppo pine forests from across Spain show drought-induced growth decline and partial recovery. Agricultural and Forest Meteorology 232: 186-194. PDF
  37. Camarero, J.J., Linares, J.C., García-Cervigón, A.I., Batllori, E., Martínez, I. and Gutiérrez, E. 2017. Back to the future: the responses of alpine treelines to climate warming are constrained by the current ecotone structure. Ecosystems 20: 683-700. PDF Supporting Information


    1. Pellizzari, E., Camarero, J.J., Gazol, A., Granda, E., Shetti, R., Wilmking, M., Moiseev, P., Pividori, M. and Carrer, M. 2016. Diverging shrub and tree growth from the Polar to the Mediterranean biomes across the European continent. Global Change Biology doi: 10.1111/gcb.13577. PDF
    2. Camarero, J.J. and Carrer, M. 2016. Bridging long-term wood functioning and nitrogen deposition to better understand changes in tree growth and forest productivity. Tree Physiology doi 10.1093/treephys/tpw111. PDF
    3. González-Casares, M., Pompa-García, M. and Camarero, J.J. 2016. Differences in climate–growth relationship indicate diverse drought tolerances among five pine species coexisting in Northwestern Mexico. Trees doi:10.1007/s00468-016-1488-0PDF
    4. Sánchez-Salguero, R., Camarero, J.J., Gutiérrez, E., González Rouco, F., Gazol, A., Sangüesa-Barreda, G., Andreu-Hayles, L., Linares, J.C. and Seftigen, K. 2016. Assessing forest vulnerability to climate warming using a process-based model of tree growth: bad prospects for rear-edges. Global Change Biology doi: 10.1111/gcb.13541. PDF
    5. Cailleret, M., Jansen, S., Robert, E.M.R., Desoto, L., Aakala, T., Antos, J.A., Beikircher, B., Bigler, C., Bugmann, H., Caccianiga, M., Čada, V., Camarero, J.J., Cherubini, P., Cochard, H., Coyea, M. R., Čufar, K., Das, A.J., Davi, H., Delzon, S., Dorman, M., Gea-Izquierdo, G., Gillner, S., Haavik, L.J., Hartmann, H., Hereş, A.-M., Hultine, K.R., Janda, P., Kane, J.M., Kharuk, V.I., Kitzberger, T., Klein, T., Kramer, K., Lens, F., Levanic, T., Linares, J.C., Lloret, F., Lobo-Do-Vale, R., Lombardi, F., López Rodríguez, R., Mäkinen, H., Mayr, S., Mészáros, I., Metsaranta, J.M., Minunno, F., Oberhuber, W., Papadopoulos, A., Peltoniemi, M., Petritan, A.M., Rohner, B., Sangüesa-Barreda, G., Sarris, D., Smith, J.M., Stan, A.B., Sterck, F., Stojanović, D.B., Suarez, M.L., Svoboda, M., Tognetti, R., Torres-Ruiz, J.M., Trotsiuk, V., Villalba, R., Vodde, F., Westwood, A.R., Wyckoff, P.H., Zafirov, N. and Martínez-Vilalta, J. 2016. A synthesis of radial growth patterns preceding tree mortality. Global Change Biology doi:10.1111/gcb.13535. PDF Supporting Information
    6. Camisón, A., Silla, F. and Camarero, J.J. 2016. Influences of the atmospheric patterns on unstable climate-growth associations of western Mediterranean forests. Dendrochronologia 40: 130142PDF
    7. Vicente-Serrano S.M., Camarero, J.J., Olano, J.M., Martín-Hernández, N., Peña-Gallardo, M., Tomás-Burguera, M., Gazol, A., Azorin-Molina, C.,  Bhuyan, U. and El Kenawy, A. 2016. Diverse relationships between forest growth and the Normalized Difference Vegetation Index at a global scale. Remote Sensing of the Environment 187: 14-29. PDF
    8. Gazol, A., Camarero, J.J., Anderegg, W.R.L. and Vicente-Serrano, S.M. 2017. Impacts of droughts on the growth resilience of Northern Hemisphere forests. Global Ecology and Biogeography 26: 166-176. PDF
    9. Marqués L., Camarero, J.J., Gazol, A. and Zavala, M.A. 2016. Drought impacts on tree growth of two pine species along an altitudinal gradient and their use as early-warning signals of potential shifts in tree species distributions. Forest Ecology and Management 381: 157-167. PDF
    10. Primicia I., Camarero, J.J., Martínez de Aragón, J., de-Miguel, S. and Bonet, J.A. 2016. Linkages between climate, seasonal wood formation and mycorrhizal mushroom yields. Agricultural and Forest Meteorology 228: 1-10. PDF
    11. Camarero, J.J., Guada, G., Sánchez-Salguero, R. and Cervantes, E. 2016. Winter drought impairs xylem phenology, anatomy and growth in Mediterranean Scots pine forests. Tree Physiology 36: 1536-1549. PDF
    12. Li, X., Rossi, S., Liang, E. and Camarero, J.J. 2016. Temperature thresholds for the onset of xylogenesis in alpine shrubs on the Tibetan PlateauTrees 30: 2091-2099. PDF
    13. Matias, L., Gonzalez-Diaz, P., Quero, J.L., Camarero, J.J., Lloret, F. and Jump, A.S. 2016. Role of geographical provenance in the response of silver fir seedlings to experimental warming and drought. Tree Physiology doi 10.1093/treephys/tpw049 PDF
    14. Buckley, H.L., Case, B.S., Vallejos, R., Camarero, J.J., Gutiérrez, E., Liang, E., Wang, Y. and Ellison A.M. 2016. Detecting ecological patterns along environmental gradients: alpine treeline ecotones. Chance 29: 10-15. PDF
    15. Anderegg W.R.L. Martinez-Vilalta, J., Cailleret, M., Camarero, J.J., Ewers, B., Galbraith, D., Gessler, A. Grote, R., Huang, C.-Y., Levick, S., Powell, T.L., Rowland, L., Sánchez-Salguero, R. and Trotsiuk, V. 2016. When a tree dies in the forest: scaling climate-driven tree mortality to ecosystem water and carbon fluxes. Ecosystems 19: 1133-1147. PDF
    16. Li, X., Camarero, J.J., Case, B., Liang, E. and Rossi, S. 2016. The onset of xylogenesis is not related to distance from the crown in Smith fir trees from south-eastern Tibetan Plateau. Canadian Journal of Forest Research 46: 885889PDF
    17. Lu, X., Huang, R., Wang, Y., Sigdel, S.R., Dawadi, B., Liang, E. and Camarero, J.J. 2016. Summer temperature drives radial growth of alpine shrub willows on the northeastern Tibetan Plateau. Arctic, Antarctic and Alpine Research 48: 461-468PDF
    18. Cailleret, M., Bigler, C. Bugmann, H., Camarero, J.J., Cufar, K., Davi, H., Mészáros, I., Minunno, F., Peltoniemi, M., Robert, E.M.R., Suarez, M.L., Tognetti, R. and Martínez‐Vilalta, J. 2016. Towards a common methodology for developing logistic tree mortality models based on ring-width data. Ecological Applications 261827-1841PDF
    19. Liang, E., Wang, Y., Piao, S., Lua, X., Camarero, J.J., Zhu, H., Zhu, L., Ellison, A.M., Ciais, P. and Peñuelas, J. 2016. Species interactions slow warming-induced upward shifts of treelines on the Tibetan Plateau. Proceedings of the National Academy of Sciences of the United States of America 113: 43804385PDF
    20. Guada, G., Camarero, J.J., Sánchez-Salguero, R. and Navarro-Cerrillo, R.M. 2016. Limited growth recovery after drought-induced forest dieback in very defoliated trees of two pine species. Frontiers in Plant Science 10.3389/fpls.2016.00418. PDF www
    21. Wang, Y., Pederson, N., Ellison, A.M., Buckley, H.L., Case, B.S., Liang, E. and Camarero, J.J. 2016. Increased stem density and competition may diminish the positive effects of warming at alpine treeline. Ecology 971668-1679. PDF
    22. Gazol, A. and Camarero, J.J. 2016. Functional diversity enhances silver fir growth resilience to an extreme drought. Journal of Ecology 10410631075. PDF  Supp Info
    23. Piper, F., Viñegla, B., Linares, J.C., Camarero, J.J., Cavieres, L.A. and Fajardo, A. 2016. Mediterranean and temperate treelines are controlled by different environmental drivers. Journal of Ecology 104691-702. PDF
    24. Pellizzari, E., Camarero, J.J., Gazol, A., Sangüesa-Barreda, G. and Carrer, M. 2016. Wood anatomy and carbon-isotope discrimination support long-term hydraulic deterioration as a major cause of drought-induced dieback. Global Change Biology 22: 21252137. PDF
    25. Wagner, F. H., Hérault, B., Bonal, D., Stahl, C., Anderson, L. O., Baker, T. R., Becker, G. S., Beeckman, H., Boanerges Souza, D., Botosso, P. C., Bowman, D. M. J. S., Bräuning, A., Brede, B., Brown, F. I., Camarero, J. J., Camargo, P. B., Cardoso, F. C. G., Carvalho, F. A., Castro, W., Chagas, R. K., Chave, J., Chidumayo, E. N., Clark, D. A., Costa, F. R. C., Couralet, C., da Silva Mauricio, P. H., Dalitz, H., de Castro, V. R., de Freitas Milani, J. E., de Oliveira, E. C., de Souza Arruda, L., Devineau, J.-L., Drew, D. M., Dünisch, O., Durigan, G., Elifuraha, E., Fedele, M., Ferreira Fedele, L., Figueiredo Filho, A., Finger, C. A. G., Franco, A. C., Freitas Júnior, J. L., Galvão, F., Gebrekirstos, A., Gliniars, R., Graça, P. M. L. D. A., Griffiths, A. D., Grogan, J., Guan, K., Homeier, J., Kanieski, M. R., Kho, L. K., Koenig, J., Kohler, S. V., Krepkowski, J., Lemos-Filho, J. P., Lieberman, D., Lieberman, M. E., Lisi, C. S., Longhi Santos, T., López Ayala, J. L., Maeda, E. E., Malhi, Y., Maria, V. R. B., Marques, M. C. M., Marques, R., Maza Chamba, H. M., Mbwambo, L., Melgaço, K. L. L., Mendivelso, H. A., Murphy, B. P., O’Brien, J. J., Oberbauer, S. F., Okada, N., Pélissier, R., Prior, L. D., Roig, F. A., Ross, M., Rossatto, D. R., Rossi, V., Rowland, L., Rutishauser, E., Santana, H., Schulze, M., Selhorst, D., Silva, W. R., Silveira, M., Spannl, S., Swaine, M. D., Toledo, J. J., Toledo, M. M., Toledo, M., Toma, T., Tomazello Filho, M., Valdez Hernández, J. I., Verbesselt, J., Vieira, S. A., Vincent, G., Volkmer de Castilho, C., Volland, F., Worbes, M., Zanon, M. L. B., and Aragão, L. E. O. C. 2016. Climate seasonality limits carbon assimilation and storage in tropical forests. Biogeosciences 13: 2537-2562, doi:10.5194/bg-2015-619 PDF www.
    26. Liu, B., Wang, Y., Zhu, H., Liang, E. and Camarero, J.J. 2016. Topography and age mediate the growth responses of Smith fir to climate warming in the southeastern Tibetan Plateau. International Journal of Biometeorology 60: 15771587. PDF
    27. Shestakova, T.A., Gutiérrez, E., Kirdyanov, A.V., Camarero, J.J., Génova, M., Knorre, A.A., Linares, J.C., Resco de Dios, V., Sánchez-Salguero, R. and Voltas, J. 2016. Forests synchronize their growth in contrasting Eurasian regions in response to climate warming. Proceedings of the National Academy of Sciences of the United States of America 113: 662667. PDF
    28. Wang, Y., Zhu, H., Liang, E. and Camarero, J.J. 2016. Impact of plot shape and size on the evaluation of treeline dynamics in the Tibetan Plateau. Trees 30: 10451056. PDF
    29. Herguido, E., Granda, E., Benavides, R., García-Cervigón, A.I., Camarero, J.J. and Valladares, F. 2016. Contrasting growth and mortality responses to climate warming of two pine species in a continental Mediterranean ecosystem. Forest Ecology and Management 363: 149158. PDF
    30. Alfaro Sanchez, R., Camarero, J.J., Sangüesa-Barreda, G. and de las Heras, J. 2016. Post-fire Aleppo pine growth, C and N isotope composition depend on site dryness. Trees 10.1007/s00468-015-1342-9. PDF
    31. Pacheco, A., Camarero, J.J. and Carrer, M. 2016. Linking wood anatomy and xylogenesis allows pinpointing of climate and drought influences on growth of coexisting conifers in continental Mediterranean climate. Tree Physiology 36: 502512. PDF
    32. Mendivelso, H.A., Camarero, J.J., Gutiérrez, E. and Castaño-Naranjo, A. 2016. Climate influences on leaf phenology, xylogenesis and radial stem changes at hourly to monthly scales in two tropical dry forests. Agricultural and Forest Meteorology 216: 20-36. PDF


  1. Cuny, H.E., Rathgeber, C.B.K., Frank, D., Fonti, P., Mäkinen, H., Prislan, P., Rossi, S., Martinez del Casillo, E., Campelo, F., Vavrčík, H., Camarero, J.J., Bryukhanova, M., Jyske, T., Gričar, J., Gryc, V., De Luis, M., Vieira, J., Čufar, K., Kirdyanov, A.V., Oberhuber, W., Treml, V., Huang, J.-G., Li, X. Swidrak, I.,  Deslauriers, A., Liang, E., Nöjd, P., Gruber, A., Nabais, C., Morin, H., Krause, C., King, G. and Fournier, M. 2015. Woody biomass production lags stem-girth increase by over one month in coniferous forests. Nature Plants 15160: 1-6. PDF
  2. Camarero, J.J., Sangüesa-Barreda, G. and Vergarechea, M. 2016. Prior height, growth, and wood anatomy differently predispose to drought-induced dieback in two Mediterranean oak species. Annals of Forest Science 73: 341351. PDF
  3. Sánchez-Salguero, R., Linares, J.C., Camarero, J.J., Madrigal-González, J., Hevia, A., Sánchez-Miranda, A., Ballesteros-Cánovas, J., Alfaro-Sánchez, R., García-Cervigón. A.I., Bigler, C. and Rigling, A. 2015. Disentangling the effects of competition and climate on individual tree growth: A retrospective and dynamic approach in Scots pineForest Ecology and Management 358: 1225. PDF
  4. Sangüesa-Barreda, G., Linares, J.C. and Camarero, J.J. 2015. Reduced growth sensitivity to climate in bark-beetle infested Aleppo pines: Connecting climatic and biotic drivers of forest dieback. Forest Ecology and Management 357: 126137. PDF
  5. Camarero, J.J., Gazol, A., Sancho-Benages, S. and Sanguesa-Barreda, G. 2015. Know your limits? Climate extremes impact the range of Scots pine in unexpected places. Annals of Botany doi: 10.1093/aob/mcv124 PDF
  6. Camarero, J.J., Gazol, A., Tardif, J.C. and Conciatori, F. 2015. Attributing forest responses to global-change drivers: limited evidence of a CO2-fertilization effect in Iberian pine growth. Journal of Biogeography 42: 2220-2233. PDF
  7. Anderegg, W.R.L., Schwalm, C., Biondi, F., Camarero, J.J., Koch, G., Litvak, M., Ogle, K., Shaw, J.D., Shevliakova, E., Williams, A.P., Wolf, A., Ziaco, E. and Pacala, S. 2015. Pervasive drought legacies in forest ecosystems and their implications for carbon cycle modelsScience 349: 528-532. PDF www 
  8. Camarero, J.J., García-Ruiz, J.M., Sangüesa-Barreda, G., Galván, J.D., Alla, A.Q., Sanjuán, Y., Beguería, S. and Gutiérrez, E. 2015. Recent and intense dynamics in a formerly static treeline. Arctic, Antarctic and Alpine Research 47: 773-783. PDF
  9. Primicia, I., Camarero, J.J., Janda, P., Čada, V., Morrissey, R.C., Trotsiuk, V., Bače, R., Teodosiu, M. and Svoboda, M. 2015. Age, competition, disturbance and elevation effects on tree and stand growth response of primary Picea abies forest to climateForest Ecology and Management 354: 7786. PDF
  10. Pompa-García, M. and Camarero, J.J. 2015. Reconstructing evaporation from pine tree rings in northern Mexico. Tree-Ring Research 71: 95-105. PDF
  11. Wang, Y., Liang, E., Ellison, A.M., Lu, X. and Camarero, J.J. 2015. Facilitation stabilizes moisture-controlled alpine juniper shrublines in the central Tibetan Plateau. Global and Planetary Change 132: 20-30. PDF
  12. Corrià-Ainslie, R., Camarero, J.J. and Toledo, M. 2015. Environmental heterogeneity and dispersal processes influence post-logging seedling establishment in a Chiquitano dry tropical forest. Forest Ecology and Management 349: 122-133. PDF
  13. Sangüesa-Barreda, G., Camarero, J.J., Oliva, J., Montes, F. and Gazol, A. 2015. Past logging, drought and pathogens interact and contribute to forest diebackAgricultural and Forest Meteorology 208: 85-94. PDF
  14. Camarero, J.J., Franquesa, M. and Sangüesa-Barreda, G. 2015. Timing of drought triggers distinct growth responses in holm oak: implications to predict warming-induced forest defoliation and growth decline. Forests 6: 1576-1597. PDF
  15. Seijo, F., Millington, J.D.A., Gray, R., Sanz, V., Lozano, J., García-Serrano, F., Sangüesa-Barreda, G. and Camarero, J.J. 2015. Forgetting fire: Traditional fire knowledge in two chestnut forest ecosystems of the Iberian Peninsula and its implications for European fire management policy. Land Use and Policy 47: 130-144. PDF
  16. Gazol, A., Camarero, J.J., Gutiérrez, E., Popa, I., Andreu-Hayles, L., Motta, R., Nola, P., Ribas, M., Sangüesa-Barreda, G., Urbinati, C. and Carrer, M. 2015. Distinct effects of climate warming on populations of silver fir (Abies alba) across Europe. Journal of Biogeography 42: 1150-1162PDF
  17. Lu, X., Camarero, J.J., Wang, Y., Liang, E. and Eckstein, D. 2015. Up to 400-year-old Rhododendron shrubs on the southeastern Tibetan Plateau: prospects for shrub-based dendrochronology. Boreas 44: 760-768PDF
  18. Sánchez-Salguero, R., Camarero, J.J., Hevia, A., Madrigal-González, J., Linares, J.C., Ballesteros-Canovas, J.A., Sánchez-Miranda, A., Alfaro-Sánchez, R., Sangüesa-Barreda, S., Galván, J.D., Gutiérrez, E., Génova, M. and Rigling, A. 2015. What drives growth of Scots pine in continental Mediterranean climates: Drought, low temperatures or both? Agricultural and Forest Meteorology 206: 151-162. PDF
  19. Vicente-Serrano, S.M., Camarero, J.J. , Zabalza, J., Sangüesa-Barreda, G., López-Moreno, J.I. and Tague, C.L. 2015. Evapotranspiration deficit controls net primary production and growth of silver fir: Implications for Circum-Mediterranean forests under forecasted warmer and drier conditionsAgricultural and Forest Meteorology 206: 45-54. PDF
  20. González de Andrés, E., Camarero, J.J. and Büntgen, U. 2015. Complex climate constraints of upper treeline formation in the Pyrenees. Trees 29: 941-952PDF
  21. Büntgen, U., Egli, S., Schneider, L., von Arx, G., Rigling, A., Camarero, J.J., Sangüesa-Barreda, G., Fischer, C.R., Oliach, D., Bonet, J.A., Colinas, C., Tegel, W., Ruiz Barbarin, J.I. and Martínez-Peña, F. 2015. Long-term irrigation effects on Spanish holm oak growth and its black truffle symbiont. Agriculture, Ecosystems and Environment 202: 148-159. PDF
  22. Camarero, J.J., Gazol, A., Sangüesa-Barreda, G., Oliva, J. and Vicente-Serrano, S. M. 2015. To die or not to die: early-warning signals of dieback in response to a severe drought. Journal of Ecology 103: 44-57. PDF
  23. Rozas, V., Camarero, J.J., Sangüesa-Barreda, G., Souto, M. and García-González, I. 2015. Summer drought and ENSO-related cloudiness distinctly drive Fagus sylvatica growth near the species rear-edge in northern Spain. Agricultural and Forest Meteorology 201: 153-164. PDF
  24. Galván, D.J., Büntgen, U., Ginzler, C., Grudd, H., Gutiérrez, E., Labuhn, I. and Camarero, J.J. 2015. Drought-induced weakening of growth-temperature associations in high-elevation Iberian pines. Global and Planetary Change 124: 95-106. PDF
  25. Camarero, J.J., Gazol, A., Galván, J.D., Sangüesa-Barreda, G. and Gutiérrez, E. 2015. Disparate effects of global-change drivers on mountain conifer forests: warming-induced growth enhancement in young trees vs. CO2 fertilization in old trees from wet sites. Global Change Biology 21: 738-749. PDF


  1. Alfaro Sánchez, R., Camarero, J.J., López-Serrano, F.R., Sánchez Salguero, R., Moya, D. and de las Heras, J. 2015. Positive coupling between growth and reproduction in young post-fire Aleppo pines depends on climate and site conditions. International Journal of Wildland Fire 24:507–517. PDF
  2. Fleta-Soriano, E., Fernández-Marín, B., Olano, J.M., Míguez, F., Molinero, J., Camarero, J.J. and García-Plazaola, J.I. 2014. Tocochromanols in wood: a potential new tool for dendrometabolomics. Tree Physiology 34: 1411-1418. PDF
  3. Heres, A., Camarero, J.J., López, B.C. and Martínez-Vilalta, J. 2014. Declining hydraulic performances and low carbon investments in tree rings predate Scots pine drought-induced mortality. Trees 28: 1737-1750. PDF
  4. Esper, J., Grossjean, J., Camarero, J.J., García-Cervigón, A.I., Olano, J.M., González-Rouco, J.F., Domínguez-Castro, F. and Büntgen, U. 2015. Atlantic and Mediterranean synoptic drivers of central Spanish juniper growth.Theoretical and Applied Climatology 12: 571-579. PDF
  5. Galván, D., Camarero, J.J., Ginzler, C. and Büntgen, U. 2014. Spatial diversity of recent trends in Mediterranean tree growth. Environmental Research Letters 9, 084001. www PDF
  6. Peguero-Pina, J.J., Sancho-Knapik, D., Barrón, E., Camarero, J.J., Vilagrosa, A. and Gil-Pelegrín, E. 2014. Morphological and physiological divergences within Quercus ilex support the existence of different ecotypes depending on climatic dryness. Annals of Botany 114: 301-313. PDF
  7. Navarro-Cerrillo, R.M., Sánchez-Salguero, R., Manzanedo, R.D., Camarero, J.J. and Fernández Cancio, A. 2014. Site and age condition the growth responses to climate and drought of relict Pinus nigra subsp. salzmanniipopulations in Southern Spain. Tree-Ring Research 70 145-155. PDF
  8. Galván, D., Camarero, J.J. and Gutiérrez, E. 2014. Seeing the trees for the forest: drivers of individual growth responses to climate in Pinus uncinata mountain forests. Journal of Ecology 102: 1244–1257. PDF
  9. Vicente-Serrano, S.M., Camarero, J.J. and Azorín-Molina, C. 2014. Diverse responses of forest growth to drought time-scales in the Northern Hemisphere. Global Ecology and Biogeography 23: 1019-1030. PDF
  10. Sangüesa-Barreda, G., Camarero, J.J., García-Martín, A., Hernández, R. and de la Riva, J. 2014. Remote-sensing and tree-ring based characterization of forest defoliation and growth loss due to the Mediterranean pine processionary moth. Forest Ecology and Management 320: 171-181. PDF
  11. Navarro-Cerrillo, R.M., Camarero, J.J., Manzanedo, R.D., Sánchez-Cuesta, R., López Quintanilla, J. and Sánchez-Salguero, R. 2014. Regeneration of Abies pinsapo within gaps created by Heterobasidion annosum-induced tree mortality in southern Spain. iForests doi:10.3832/ifor0961-007 PDF
  12. González de Andrés, E., Camarero, J.J., Martínez, I. and Coll, Ll. 2014. Uncoupled spatiotemporal patterns of seed dispersal and regeneration in Pyrenean silver fir populations. Forest Ecology and Management 319: 18-28. PDF
  13. Camarero, J.J., Rozas, V. and Olano, J.M. 2014. Minimum wood density of Juniperus thurifera is a robust proxy of spring water availability in a continental Mediterranean climate. Journal of Biogeography 41: 1105-1114PDF Appendix
  14. Nuche, P., Komac, B., Camarero, J.J. and Alados, C.L. 2014. Developmental instability as an index of adaptation to drought stress in a Mediterranean oak. Ecological Indicators 40: 68-75. PDF
  15. Mendivelso, H.A., Camarero, J.J., Gutiérrez, E. and Zuidema, P.A. 2014. Time-dependent effects of climate and drought on tree growth in a Neotropical dry forest: Short-term tolerance vs. long-term sensitivity. Agricultural and Forest Meteorology 188: 13-23. PDF
  16. Büntgen, U., Tegel, W., Kaplan, J.O., Schaub, M., Hagedorn, F., Bürgi, M., Brázdil, R., Helle, G., Carrer, M., Heussner, K.-U., Hofmann, J., Kontic, R., Kyncl, T., Kyncl, J., Camarero, J.J., Tinner, W., Esper, J. and Liebhold, A.2014. Placing unprecedented recent fir growth in a European-wide and Holocene-long context. Frontiers in Ecology and the Environment 12: 100-106PDF
  17. Granda, E., Rossatto, D.R., Camarero, J.J., Voltas, J. and Valladares, F. 2014. Growth and carbon isotopes of Mediterranean trees reveal contrasting responses to increased carbon dioxide and drought. Oecologia 174: 307-317. PDF 


  1. Sánchez-Salguero, R., Camarero, J.J., Dobbertin, M., Fernández-Cancio, A., Vilà-Cabrera, A., Manzanedo, R.D., Zavala, M.A. and Navarro-Cerillo, R.M. 2013. Contrasting vulnerability and resilience to drought-induced decline of densely planted vs. natural rear-edge Pinus nigra forests. Forest Ecology and Management 310: 956-967. PDF 
  2. Mendivelso, H.A., Camarero, J.J., Royo Obregón, O., Gutiérrez, E. and Toledo, M. 2013. Differential growth responses to water balance of coexisting deciduous tree species are linked to wood density in a Bolivian tropical dry forest. Plos ONE 8: e73855 doi:10.1371/journal.pone.0073855. www PDF
  3. Amanzadeh, B., Sagheb-Talebi, K., Sotoudeh Foumani, B., Fadaie, F., Camarero, J.J. and Linares, J.C. 2013. Spatial distribution and volume of dead wood in unmanaged caspian beech (Fagus orientalis) forests from northern Iran. Forests 4: 751-765. doi:10.3390/f4040751. www PDF
  4. Camarero, J.J., Manzanedo, R.D., Sánchez-Salguero, R. and Navarro-Cerrillo, R. 2013. Growth response to climate and drought change along an aridity gradient in the southernmost Pinus nigra relict forests. Annals of Forest Science 70: 769-780PDF
  5. Touchan, R., Meko, D.M., Ballesteros, J.A., Sánchez-Salguero, R., Camarero, J.J., Kerchouche, D., Muntán, E., Khabcheche, M., Blanco, J.A., Rodríguez Morata, C., Garófano-Gómez, V., Martín, L.A., Alfaro-Sánchez, R., Garah, K., Hevia, A., Madrigal-González, J., Sánchez-Miranda, A., Shestakova, T.A. and Tabakova, M. 2013. Dendrochronology course in Valsaín Forest, Segovia, Spain. Tree-Ring Research 69: 93-100. PDF
  6. Pardo, I., Camarero, J.J., Gutiérrez, E. and García, M.B. 2013. Uncoupled changes in tree cover and field layer vegetation at two Pyrenean treelines over 11 years. Plant Ecology and Diversity 6: 355-364. PDF
  7. Morán-Tejeda, E., Ceglar, A., Medved-Cvikl, B., Vicente-Serrano, S.-M., López-Moreno, J.I., González-Hidalgo, J.C., Revuelto, J., Lorenzo-Lacruz, J., Camarero, J.J. and Pasho, E. 2013. Assessing the capability of multi-scale drought datasets to quantify drought severity and to identify drought impacts: an example in the Ebro Basin. International Journal of Climatology 33: 1884-1897. PDF
  8. Büntgen, U., Martínez-Peña, F., Aldea, J., Rigling, A., Fischer, E.M., Camarero, J.J., Hayes, M.J., Fatton, V. and Egli, S. 2013. Declining pine growth in Central Spain coincides with increasing diurnal temperature range since the 1970s. Global and Planetary Change 107: 177-185. PDF
  9. Alla, A.Q., Camarero, J.J., Palacio. S, and Montserrat-Martí, G. 2013. Revisiting the fate of buds: size and position drive bud mortality and bursting in two coexisting Mediterranean Quercus species with contrasting leaf habit. Trees 27: 1375-1386. PDF
  10. García-Cervigón, A.I., Gazol, A., Sanz, V., Camarero, J.J. and Olano, J.M. 2013. Intraspecific competition replaces interspecific facilitation as abiotic stress decreases: The shifting nature of plant–plant interactions. Perspectives in Plant Ecology, Evolution and Systematics 15: 226-236PDF
  11. Sangüesa-Barreda, G., Linares, J.C. and Camarero, J.J. 2013. Drought and mistletoe reduce growth and water-use efficiency of Scots pine. Forest Ecology and Management 296: 64-73. PDF
  12. Voltas, J., Camarero, J.J., Carulla, D., Aguilera, M., Oriz, A. and Ferrio, J.P. 2013. A retrospective, dual-isotope approach reveals individual predispositions to winter-drought induced tree dieback in the southernmost distribution limit of Scots pine. Plant, Cell and Environment 36: 1435-1448. PDF
  13. Granda, E., Camarero, J.J., Jimeno, T.E., Martínez-Fernández, J. and Valladares, F. 2013. Intensity and timing of warming and drought differentially affect growth patterns of co-occurring Mediterranean tree species. European Journal of Forest Research 132: 469-480. PDF
  14. Primicia, I., Camarero, J.J., Imbert, J.B. and Castillo, F. J. 2013. Effects of thinning and canopy type on growth dynamics of Pinus sylvestris: inter-annual variations and intra-annual interactions with microclimate. European Journal of Forest Research 132: 121-135. PDF
  15. Linares, J.C., Taïqui, L., Sangüesa-Barreda, G., Seco, J.I. and Camarero, J.J. 2013. Age-related drought sensitivity of Atlas cedar (Cedrus atlantica) in the Moroccan Middle Atlas forests. Dendrochronologia 31: 88-96. PDF
  16. Camarero, J.J., Palacio, S. and Montserrat-Martí, G. 2013. Contrasting seasonal overlaps between primary and secondary growth are linked to wood anatomy in Mediterranean sub-shrubs. Plant Biology 15: 798-807. PDF
  17. Vicente-Serrano, S.M., Gouveia, C., Camarero, J.J., Beguería, S., Trigo, R., López-Moreno, J.I., Azorín-Molina, C., Pasho, E., Lorenzo-Lacruz, J., Revuelto, J., Morán-Tejeda, E. and Sanchez-Lorenzo, A. 2013. Response of vegetation to drought time-scales across global land biomes. Proceedings of the National Academy of Sciences of the United States of America 110: 52-57. PDF


  1. Linares, J.C. and Camarero, J.J. 2012. Silver fir defoliation likelihood is related to negative growth trends and high warming sensitivity at their southernmost distribution limit. ISRN Forestry doi: 10.5402/2012/437690. PDF
  2. Büntgen, U., Egli, E., Camarero, J.J., Fischer, E.M., Stobbe, U., Kauserud, H., Tegel, W., Sproll, L., Stenseth, N.C. 2012. Drought-induced decline in Mediterranean truffle harvest. Nature Climate Change 2: 827-829. PDF
  3. Alla, A.Q., Camarero, J.J. and Montserrat-Martí, G. 2012. Seasonal and inter-annual variability of bud development as related to climate in two co-existing Mediterranean Quercus species. Annals of Botany 111: 261-270. PDF
  4. Coll, Ll., Camarero, J.J. and Martínez de Aragón, J. 2012. Fine root seasonal dynamics, plasticity, and mycorrhization in 2 coexisting Mediterranean oaks with contrasting aboveground phenology. Écoscience 19: 238-245. PDF
  5. Vicente-Serrano, S.M., Beguería, S., Lorenzo-Lacruz, J., Camarero, J.J., López-Moreno, J.I., Azorin-Molina, C., Revuelto, J., Morán-Tejeda, E. and Sánchez-Lorenzo, A. 2012. Performance of drought indices for ecological, agricultural, and hydrological applications. Earth Interactions 16:1-27. PDF
  6. Wang, Y., Camarero, J.J., Luo, T. and Liang, E. 2012. Spatial patterns of Smith fir alpine treelines on the south-eastern Tibetan Plateau support that contingent local conditions drive recent treeline patterns. Plant Ecology and Diversity 5: 311-321. PDF
  7. Gazol, A. and Camarero, J.J. 2012. Mediterranean dwarf shrubs and coexisting trees present different radial-growth synchronies and responses to climate. Plant Ecology 213: 1687-1698. PDF
  8. García-Cervigón, A.I., Olano, J.M., Eugenio, M. and Camarero, J.J. 2012. Arboreal and prostrate conifers coexisting in Mediterranean high mountains differ in their climatic responses. Dendrochronologia 30: 279-286. PDF
  9. Palacio, S., Hernández, R., Maestro-Martínez, M. and Camarero, J.J. 2012. Fast replenishment of initial carbon stores after defoliation by the pine processionary moth and its relationship to the re-growth ability of trees. Trees 26: 1627-1640. PDF
  10. Morán-Tejeda, E., Ceglar A., Medved-Cvikl, B., Vicente-Serrano, S.M., López-Moreno, J.I., González-Hidalgo, J.C., Revuelto, J., Lorenzo-Lacruz, J., Camarero, J.J. and Pasho, E. 2012. Assessing the capability of multi-scale drought datasets to quantify drought severity and to identify drought impacts: an example in the Ebro Basin. International Journal of Climatology 33: 1884-1897. PDF
  11. Pasho, E., Camarero, J.J. and Vicente-Serrano, S.M. 2012. Climatic impacts and drought control of radial growth and seasonal wood formation in Pinus halepensis. Trees 26: 1875-1886. PDF
  12. Camarero, J.J., Olano, J.M., Arroyo Alfaro, S.J., Fernández-Marín, B., Becerril, J.M. and García-Plazaola, J.I. 2012. Photoprotection mechanisms in Quercus ilex under contrasting climatic conditions. Flora 207:557-564. PDF
  13. Gazol, A. and Camarero, J.J. 2012. The performance of Mediterranean subshrubs depends more on microsite than on regional climate conditions. Journal of Vegetation Science 23: 1062-1070. PDF
  14. Galván, J.D., Camarero, J.J., Sangüesa-Barreda, G., Alla, A.Q. and Gutiérrez, E. 2012. Sapwood area drives growth in mountain conifer forests. Journal of Ecology 100: 1233-1244. PDF
  15. Pasho, E., Camarero, J.J., de Luis, M. and Vicente-Serrano, S.M. 2012. Factors driving growth responses to drought in Mediterranean forests. European Journal of Forest Research 131: 1797-1807. PDF
  16. Sangüesa-Barreda, G., Linares, J.C. and Camarero, J.J. 2012. Mistletoe effects on Scots pine decline following drought events: insights from within-tree spatial patterns, growth and carbohydrates. Tree Physiology 32:585-598.PDF
  17. Oliva, J., Camarero, J.J. and Stenlid, J. 2012. Understanding the role of sapwood loss and reaction zone formation on radial growth of Norway spruce (Picea abies) trees decayed by Heterobasidion annosum s.l. Forest Ecology and Management 274: 201-209. PDF
  18. Kouba, Y., Camarero, J.J. and Alados, C.L. 2012. Roles of land-use and climate change on the establishment and regeneration dynamics of Mediterranean semi-deciduous oak forests. Forest Ecology and Management 274: 143-150. PDF
  19. Gimeno, T.E., Camarero, J.J., Granda, E., Pías, B. and Valladares, F. 2012. Enhanced growth of Juniperus thurifera under a warmer climate is explained by a positive carbon gain under cold and drought. Tree Physiology 32:326-336. PDF
  20. Alla, A.Q. and Camarero, J.J. 2012. Contrasting responses of radial growth and wood anatomy to climate in a Mediterranean ring-porous oak: implications for its future persistence or why the variance matters more than the mean. European Journal of Forest Research 131: 1537-1550. PDF
  21. Martínez I., González-Taboada F., Wiegand T., Camarero, J.J. and Gutiérrez, E. 2012. Dispersal limitation and spatial scale affect model based projections of Pinus uncinata response to climate change in the Pyrenees. Global Change Biology 18: 1714-1724. PDF
  22. Linares, J.C. and Camarero, J.J. 2012. From pattern to process: linking intrinsic water-use efficiency to drought-induced forest decline. Global Change Biology 18: 1000-1015. PDF
  23. Alla, A.Q., Camarero, J.J., Maestro-Martínez, M. and Montserrat-Martí, G. 2012. Acorn production is linked to secondary growth but not to declining carbohydrate concentrations in current-year shoots of two oak species. Trees 26: 841-850. PDF
  24. Sánchez-Salguero, R., Navarro-Cerillo, R.M., Camarero, J.J. and Fernández-Cancio, A. 2012. Selective drought-induced decline of pine species in southeastern Spain. Climatic Change 113: 767-785. PDF
  25. DeSoto, L., Camarero, J.J., Olano, J.M. and Rozas, V. 2012. Geographically structured and temporally unstable growth responses of Juniperus thurifera to recent climate variability in the Iberian Peninsula. European Journal of Forest Research 131: 905-917. PDF
  26. Linares, J.C. and Camarero, J.J. 2012. Growth patterns and sensitivity to climate predict silver fir decline in the Spanish Pyrenees. European Journal of Forest Research 131: 1001-1012. PDF


  1. Komac, B., Alados, C.L. and Camarero, J.J. 2011. Influence of topography on the colonization of subalpine grasslands by the thorny cushion dwarf Echinospartum horridum.  Arctic, Antarctic and Alpine Research 43: 601-611. PDF
  2. Pasho, E., Camarero, J.J., de Luis, M. and Vicente-Serrano, S.M. 2011. Impacts of drought at different time scales on forest growth across a wide climatic gradient in north-eastern Spain. Agricultural and Forest Meteorology 151: 1800-1811. PDF
  3. Linares, J.C., Taïqui, L. and Camarero, J.J. 2011. Increasing drought sensitivity and decline of Atlas cedar (Cedrus atlantica) in the Moroccan Middle Atlas forests. Forests 2: 777-796. doi:10.3390/f2030777. www PDF
  4. Alla, A.Q., Camarero, J.J., Rivera, P. and Montserrat-Martí, G. 2011. Variant allometric scaling relationships between bud size and secondary shoot growth in Quercus faginea: implications for the climatic modulation of canopy growth. Annals of Forest Science 68: 1245-1254. PDF
  5. Camarero, J.J., Bigler, C., Linares, J.C. and Gil-Pelegrín, E. 2011. Synergistic effects of past historical logging and drought on the decline of Pyrenean silver fir forests. Forest Ecology and Management 262: 759-769. PDF
  6. Martínez,I., Wiegand, T., Camarero, J.J., Batllori, E. and Gutiérrez, E. 2011. Disentangling the formation of contrasting tree-line physiognomies combining model selection and Bayesian parameterization for simulation models. The American Naturalist 177: E136-E152. PDF
  7. Pasho, E., Camarero, J.J., de Luis, M. and Vicente-Serrano, S.M. 2011. Spatial variability in large-scale and regional atmospheric drivers of Pinus halepensis growth in eastern Spain. Agricultural and Forest Meteorology151: 1106-1119. PDF
  8. Sanz, R., Pulido, F. and Camarero, J.J. 2011. Boreal trees in the Mediterranean: recruitment of downy birch (Betula alba) at its southern range limit. Annals of Forest Science 68: 793-802. PDF
  9. Gutiérrez, E., Campelo, F., Camarero, J.J., Ribas, M., Muntán, E., Nabais, C. and Freitas, H. 2011. Climate controls act at different scales on the seasonal pattern of Quercus ilex L. stem radial increments in NE Spain. Trees25: 637-646. PDF



  1. Sánchez-Salguero, R., Navarro-Cerillo, R.M., Camarero, J.J. and Fernández-Cancio, A. 2010. Drought-induced growth decline of Aleppo and maritime pine forests in south-eastern Spain. Forest Systems 19: 458-469. PDF
  2. Batllori, E., Camarero, J.J. and Gutiérrez, E. 2010. Current regeneration patterns at the tree line in the Pyrenees indicate similar recruitment processes irrespective of the past disturbance regime. Journal of Biogeography37: 1938-1950. PDF
  3. Camarero, J.J., Albuixech, J., López-Lozano, R., Casterad, M.A. and Montserrat-Martí, G. 2010. An increase in canopy cover leads to masting in Quercus ilex. Trees 24: 909-918. PDF
  4. Linares J.C., Camarero, J.J., Bowker, M.A., Ochoa, V. and Carreira, J.A. 2010. Stand-structural effects on Heterobasidion abietinum-related mortality following drought events in Abies pinsapo. Oecologia 164: 1107-1119. PDF
  5. Linares J.C., Camarero, J.J. and Carreira, J.A. 2010. Competition modulates the adaptation capacity of forests to climatic stress: insights from recent growth decline and death in relict stands of the Mediterranean fir Abies pinsapo. Journal of Ecology 185: 471-480. PDF
  6. Camarero, J.J., Olano, J.M. and Parras, A. 2010. Plastic bimodal xylogenesis in conifers from continental Mediterranean climates. New Phytologist 185: 471-480. PDF



  1. Linares J.C., Camarero, J.J. and Carreira, J.A. 2009. Plastic responses of Abies pinsapo xylogenesis to drought and competition. Tree Physiology 29: 1525-1536. PDF
  2. Linares J.C., Delgado-Huertas, A. Camarero, J.J., Merino, J. and Carreira, J.A. 2009. Competition and drought limit the response of water-use efficiency to rising atmospheric carbon dioxide in the Mediterranean fir Abies pinsapo. Oecologia 161: 611-624. PDF
  3. Linares J.C., Camarero, J.J. and Carreira, J.A. 2009. Interacting effects of climate and forest-cover changes on mortality and growth of the southernmost European fir forests. Global Ecology and Biogeography 18: 485-497. PDF
  4. Batllori E., Camarero, J.J., Ninot J.M. and Gutiérrez, E. 2009. Seedling recruitment, survival and facilitation in alpine Pinus uncinata treeline ecotones. Implications and potential responses to climate warming. Global Ecology and Biogeography 18: 460-472. PDF
  5. Montserrat-Martí, G., Camarero, J. J., Palacio, S., Pérez-Rontomé, C., Milla, R., Albuixech, J. and Maestro, M. 2009. Summer-drought constrains the phenology and growth of two co-existing Mediterranean oaks with contrasting leaf habit: implications for their persistence and reproduction. Trees 23: 787-799. PDF



  1. Palacio, S., Milla, R., Albuixech, J., Pérez-Rontomé, C., Camarero, J. J., Maestro, M. and Montserrat-Martí, G. 2008. Seasonal variability of dry matter content and its relationship to shoot growth and non-structural carbohydrates. New Phytologist 180: 133-142. PDF



  1. Martínez-Alonso, C., Valladares, F., Camarero, J. J., López Arias, M., Serrano, M. and Rodriguez, J. A. 2007. The uncoupling of secondary growth, cone and litter production by intradecadal climatic variability in a Mediterranean Scots Pine Forest. Forest Ecology and Management 253: 19-29. PDF
  2. Camarero, J.J. and Gutiérrez, E. 2007. Response of Pinus uncinata recruitment to climate warming and changes in grazing pressure in an isolated population of the Iberian System (NE Spain). Arctic, Antarctic and Alpine Research 39: 210-217. PDF
  3. Peguero-Pina, J.J., Camarero, J.J., Abadía, A., Martín, E., González-Cascón, R., Morales, F. and Gil-Pelegrín, E. 2007. Physiological performance of silver-fir (Abies alba Mill.) populations under contrasting climates near the south-western distribution limit of the species. Flora 202: 226-236. PDF
  4. Andreu, L., Gutiérrez, E., Macias, M., Ribas, M., Bosch, O. and Camarero, J.J. 2007. Climate increases regional tree-growth variability in Iberian pine forests. Global Change Biology 13: 1-12. PDF



  1. Macias, M., Andreu, L., Bosch, O., Camarero, J.J. and Gutiérrez, E. 2006. Increasing aridity is enhancing silver fir (Abies alba Mill.) water stress in its south-western distribution limit. Climatic Change 79: 289-313. PDF
  2. Esteso-Martínez, J., Valladares, F., Camarero, J.J. and Gil-Pelegrín, E. 2006. Crown architecture and leaf habit are associated with intrinsically different light-harvesting efficiencies in Quercus seedlings from contrasting environments. Annals of Forest Science 63: 511-518. PDF
  3. Esteso-Martínez, J., Camarero, J.J. and Gil-Pelegrín, E. 2006. Competitive effects of herbs on Quercus faginea seedlings inferred from vulnerability curves and spatial-pattern analyses in a Mediterranean stand (Iberian System, northeast Spain). Écoscience 13: 378-387. PDF
  4. Wiegand, T., Camarero, J.J., Rüger, N. and Gutiérrez, E. 2006. Abrupt population changes in treeline ecotones along smooth gradients. Journal of Ecology 94: 880-892. PDF
  5. Camarero, J.J., Gutiérrez, E. and Fortin, M.-J. 2006. Spatial patterns of plant richness across treeline ecotones in the Pyrenees reveal different locations for richness and tree-cover boundaries. Global Ecology and Biogeography 15: 182-191. PDF
  6. Solla, A. and Camarero, J.J. 2006. Spatial patterns and environmental factors affecting the presence of Melampsorella caryophyllacearum infections in an Abies alba forest in NE Spain. Forest Pathology 36: 165-175. PDF
  7. Solla, A., Sánchez-Miranda, A. and Camarero, J.J. 2006. Radial-growth and wood anatomical changes in Abies alba infected by Melampsorella caryophyllacearum: a dendroecological assessment of fungal damage. Annals of Forest Science 63: 1-8. PDF
  8. Corcuera, L., Camarero, J.J., Sisó, S. and Gil-Pelegrín, E. 2006. Radial-growth and wood-anatomical changes in overaged Quercus pyrenaica coppice stands: functional responses in a new Mediterranean landscape. Trees20: 91-98. PDF



  1. Camarero, J.J., Gutiérrez, E., Fortin, M.-J. and Ribbens, E. 2005. Spatial patterns of tree recruitment in a relict population of Pinus uncinata: densification through stratified-diffusion. Journal of Biogeography 32: 1979-1992. PDF



  1. Himrane, H., Camarero, J. J. and Gil-Pelegrín, E. 2004. Morphological and ecophysiological variation of the hybrid oak Quercus subpyrenaica (Q. faginea x Q. pubescens). Trees 18: 566-575. PDF
  2. Corcuera, L., Camarero, J. J. and Gil-Pelegrín, E. 2004. Effects of a severe drought on growth and wood-anatomical properties of Quercus faginea. IAWA Journal 25: 185-204. PDF
  3. Camarero, J. J. and Gutiérrez, E. 2004. Pace and pattern of recent treeline dynamics: response of ecotones to climatic variability in the Spanish Pyrenees. Climatic Change 63: 181-200. PDF
  4. Corcuera, L., Camarero, J. J. and Gil-Pelegrín, E. 2004. Effects of a severe drought on Quercus ilex radial growth and xylem anatomy. Trees 18: 83-92. PDF



  1. Tardif, J., Camarero, J. J., Ribas, M. and Gutiérrez, E. 2003. Spatiotemporal variability in tree ring growth in the Central Pyrenees: climatic and site influences. Ecological Monographs 73: 241-257. PDF
  2. Camarero, J. J., Martín-Bernal, E. and Gil-Pelegrín, E. 2003. The impact of a needleminer (Epinotia subsequana) outbreak on radial growth of silver fir (Abies alba) in the Aragón Pyrenees: A dendrochronological assessment. Dendrochronologia 21: 1-10. PDF



  1. Corcuera, L., Camarero, J. J. and Gil-Pelegrín, E. 2002. Functional groups in Quercus species derived from the analysis of pressure-volume curves. Trees 16: 465-472. PDF
  2. Camarero, J. J. and Gutiérrez, E. 2002. Plant species distribution across two contrasting treeline ecotones in the Spanish Pyrenees. Plant Ecology 162: 247-257. PDF
  3. Camarero, J. J. and Gutiérrez, E. 2002. A multivariate approach to the study of the spatial structure of treeline ecotones. Community Ecology 3: 9-18. PDF



  1. Sisó, S., Camarero, J. J. and Gil-Pelegrín, E. 2001. Relationship between hydraulic resistance and leaf morphology in broadleaf Quercus species: a new interpretation of leaf lobation. Trees 15: 341-345. PDF



  1. Camarero, J. J., Gutiérrez, E. and Fortin, M.-J. 2000. Spatial pattern of subalpine forest-alpine grassland ecotones in the Spanish Central Pyrenees. Forest Ecology and Management 134: 1-16. PDF
  2. Camarero, J. J., Gutiérrez, E. and Fortin, M.-J. 2000. Boundary detection in altitudinal treeline ecotones in the Spanish Central Pyrenees. Arctic and Alpine Research 32: 117-126. PDF



  1. Camarero, J. J. and Gutiérrez, E. 1999. Structure and recent recruitment at alpine forest-pasture ecotones in the Spanish Central Pyrenees. Écoscience 6: 451-464. PDF



  1. Camarero, J. J., Guerrero-Campo, J. and Gutiérrez, E. 1998. Tree-ring structure and growth of Pinus uncinata Ram. and Pinus sylvestris L. in the Central Spanish Pyrenees. Arctic and Alpine Research 30: 1-10. PDF

Book Chapters

  1. Vicente-Serrano, S.M., Beguería, S. and Camarero, J.J. 2017. Drought severity in a changing climate. In: S. Eslamian (ed.), Handbook of Drought and Water Scarcity, Vol. 2: Environmental Impacts and Analysis of Drought and Water Scarcity, pp. 279-303Taylor and Francis. PDF
  2. Camarero, J.J., Linares, J.C., Sangüesa-Barreda, G., Sánchez-Salguero, R., Gazol, A., Navarro-Cerrillo, R.M. and Carreira, J.A. 2017. The Multiple Causes of Forest Decline in Spain: Drought, Historical Logging, Competition and Biotic Stressors. In: M.M. Amoroso, L.D. Daniels, P.J. Backer and J.J. Camarero (eds.), Dendroecology: Tree-ring analyses applied to ecological studies, pp. 307-323. Springer International Publishing AG. PDF
  3. Camarero, J.J. 2017. The Multiple Factors Explaining Decline in Mountain Forests: Historical Logging and Warming-Related Drought Stress is Causing Silver-Fir Dieback in the Aragón Pyrenees. In: J. Catalan, J.M. Ninot, M. Aniz and M. Mercè (eds.), High Mountain Conservation in a Changing World, Advances in Global Change Research 62, pp. 131-154Springer. PDF
  4. Camarero, J.J., Garcés-Pastor, S., Gutiérrez, E., Rull, V., Vegas-Vilarrubia, T., Canellas-Bolta, N., Sangüesa Barreda, G., Trapote, M.C., Clavaguera, A.,
    Calero, M.A., Galván Candela, J.D., Sánchez Salguero, R., Giralt, S. y Valero-Garcés, B. 2016. Reconstruyendo la historia de los bsoques pirenaicos. En: Proyectos de investigación en parques nacionales: convocatoria 2011-2014, pp. 141-156. PDF
  5. Montes, F., Gómez, C., Camarero, J.J., Coll, Ll. y Sánchez-González, M. 2016. Pasado, presente y futuro de los bosques de montaña: seguimiento y modelización de los efectos del cambio climático y la gestión sobre la dinámica forestal. En: La investigació al Parc Nacional d’Aigüestortes i Estany de Sant Maurici, X Jornades sobre Recerca al Parc Nacional d’Aigüestortes i Estany de Sant Maurici, Espot (Pallars Sobirà, Catalunya), 14-16 d’octubre de 2015, pp. 107-114. PDF
  6. Garcés-Pastor, S., Vegas-Vilarrúbia, T., Cañellas-Boltà, N., Rull, V., Calero, M.A., Giralt, S., Valero-Garcés, B., Gutiérrez, E. y Camarero, J.J. 2016. Canvis limnològics i paisatgístics durant el darrer mil·lenni a l’Estany de Sant Maurici: Estudi Multiproxy. En: La investigació al Parc Nacional d’Aigüestortes i Estany de Sant Maurici, X Jornades sobre Recerca al Parc Nacional d’Aigüestortes i Estany de Sant Maurici, Espot (Pallars Sobirà, Catalunya), 14-16 d’octubre de 2015, pp. 33-42. PDF
  7. Camarero, J.J., Gutiérrez, E., Sangüesa Barreda, G. y Galván, D. 2016. El Parque Nacional de Aigüestortes i Estany de Sant Maurici como santuario
    de bosques y árboles viejos de pino negro. En: La investigació al Parc Nacional d’Aigüestortes i Estany de Sant Maurici, X Jornades sobre Recerca al Parc Nacional d’Aigüestortes i Estany de Sant Maurici, Espot (Pallars Sobirà, Catalunya), 14-16 d’octubre de 2015, pp. 89-95. PDF
  8. Linares, J.C., Tíscar, P.A., Camarero, J.J., Sangüesa Barreda, G., Domínguez-Clavijo, M. y Carreira, J.A. 2015. Efectos del cambio climático sobre el crecimiento de Abies pinsapo y Pinus nigra salzmannii en el sur de la península ibérica: Tendencias pasadas, presentes y futuras. En: Los Bosques y la Biodiversidad frente al Cambio Climático: Impactos, Vulnerabilidad y Adaptación en EspañaHerrero, A. y Zavala M.A., eds. MAGRAMA, Madrid, España, pp. 255-269. PDF
  9. Sangüesa-Barreda, G., Camarero, J. J. y Linares, J.C. 2015. La sequía y el muérdago actúan como factores de estrés combinados en el decaimiento de bosques  de pino silvestre. En: Los Bosques y la Biodiversidad frente al Cambio Climático: Impactos, Vulnerabilidad y Adaptación en EspañaHerrero, A. y Zavala M.A., eds. MAGRAMA, Madrid, España, pp. 303-311. PDF
  10. Camarero, J. J., Sangüesa-Barreda, G. y Linares, J.C. 2015. El decaimiento de abetares pirenaicos como paradigma de vulnerabilidad de los bosques ante el cambio climático. En: Los Bosques y la Biodiversidad frente al Cambio Climático: Impactos, Vulnerabilidad y Adaptación en EspañaHerrero, A. y Zavala M.A., eds. MAGRAMA, Madrid, España, pp. 243-253. PDF
  11. Camarero, J. J. 2014. La detección de fronteras ecológicas y la descripción cuantitativa de ecotonos como herramientas para la delimitación de los páramos. En: Visión Socioecosistémica de los Páramos y la Alta Montaña Colombiana: Memorias del Proceso de Definición de Criterios para la Delimitación de Páramos. Ed. Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Bogotá D.C., Colombia, pp. 103-110. PDF
  12. Camarero, J. J., Galván Candela, J.D., Sangüesa-Barreda, G., Alla, A.Q., Chaparro Mendivelso, H.A.y Gutiérrez Merino, E. 2012. Determinantes de las respuestas individuales de los árboles al clima e implicaciones para las reconstrucciones dendroclimáticas: un caso de estudio en bosques de montaña de Pinus uncinata. En: Proyectos de Investigación en Parques Nacionales: 2008-2011, OAPN-MAGRAMA, pp. 235-267.  PDF
  13. Batllori, E., Camarero, J. J. and Gutiérrez, E. 2012. Climatic Drivers of Tree Growth and Recent Recruitment at the Pyrenean Alpine Tree Line Ecotone. In: Ecotones Between Forest and Grassland, Myster, R.W. (Ed.), Springer, pp. 247-269. PDF
  14. Camarero, J. J. 2011. Direct and Indirect Effects of the North Atlantic Oscillation on Tree Growth and Forest Decline in Northeastern Spain. In: Hydrological, Socioeconomic and Ecological Impacts of the North Atlantic Oscillation in the Mediterranean Region, Vicente-Serrano S.M. & Trigo, R.M. (Eds.), Advances in Global Change Research 46, Springer, pp. 129-152. PDF
  15. Linares, J.C., Tíscar, P.A., Camarero, J. J. , Taïqui, L., Viñegla, B., Seco, J.A., Merino, J. & Carreira, J.A. 2011. Tree growth decline on relict Western-Mediterranean mountain forests: causes and impacts. In: Forest Decline: Causes and Impacts, Chapter 4, Ed. J.A. Jenkins. Nova Science Publishers, ISBN 978-1-61470-002-9. PDF
  16. Gil-Pelegrín, E., Peguero-Pina, J.J., Camarero, J.J. , Fernández-Cancio & Navarro-Cerrillo, R. 2008. Drought and Forest Decline in the Iberian Peninsula: A Simple Explanation for a Complex Phenomenom? In: Droughts: Causes, Effects and Predictions, Ed. J.M. Sánchez, Chapter 1, pp. 27-68. Nova Science Publishers, ISBN 978-1-61470-002-9. PDF
  17. Ninot, J.M., Grau, O., Batllori, E., Camarero, J.J. & Carrillo, E. 2008. Winter drought impairs pine regeneration at the Pyrenean treeline. In: Droughts: Causes, Effects and Predictions, Ed. J.M. Sánchez, pp. 11-18. Nova Science Publishers, ISBN 978-1-60456-285-9. PDF
  18. Gutiérrez, E., Ribas, M., Planells, O., Camarero, J.J. i Tardif, J. 2003. Resposta dels arbres i dels boscos al canvi climàtic. Un tast de canvi global, Cèlia Marrasé i J.E. Llebot, (eds.). Treballs de la SCB 54: 85-110. PDF

Publications in Spanish

  1. Camarero, J.J., Gutiérrez, E. Sangüesa-Barreda, G. y Galván, J.D. 2016. El Parque Nacional de Ordesa y Monte Perdido alberga pinos negros viejos. Jornadas de Investigación P.N. Ordesa y Monte Perdido, Torla, Torla, Huesca, pp. 21-26. PDF
  2. Calero, M.A., Valero-Garcés, B.L., Rull, V., Vegas-Vilarrúbia, T., Garcés-Pastor, S., López-Vila, J. y Camarero, J.J. 2016. El registro sedimentario del lago Sant Maurici (Pirineos Centrales). Geogaceta 59: 11-14. PDF
  3. Pompa-García y Camarero, J.J. 2015. Potencial dendroclimático de la madera temprana y tardía de Pinus cooperi Blanco. Agrociencia 49: 177-187. PDF
  4. Ortega Martínez, M. y Camarero, J.J. 2016. La Memoria de los Árboles: La información del territorio contenida en los anillos de crecimiento de la madera. Naturaleza Aragonesa 33: 47-53. PDF
  5. Mendivelso, H.A., Camarero, J.J. y Gutiérrez, E. 2016. Dendrocronología en bosques neotropicales secos: métodos, avances y aplicaciones. Ecosistemas 25: 66-75. PDF
  6. García, M.B., Alados, C.L., Antor, R., Benito Alonso, J.L., Camarero, J.J., Carmena, F., Errea,  P., Fillat, F., García-González, R., García-Ruiz, J.M., Gartzia, M., Gómez García, D., Gómez, I., González-Sampériz, P., Gutiérrez, E., Jiménez, J.J., López-Moreno, J.I., Mata, P., Moreno, A., Montserrat, P., Nuche, P., Pardo, I., Revuelto, J., Rieradevall, M., Sáiz, H., Tejero, P., Vicente-Serrano, S., Villagrasa, E., Villar, L. y Valero-Garcés, B. 2016. Integrando escalas y métodos LTER para comprender la dinámica global de un espacio protegido de montaña: el Parque Nacional de Ordesa y Monte Perdido. Ecosistemas 25: 19-30. PDF
  7. Sangüesa Barreda, G., Camarero, J.J., Linares, J.C., Hernández, R., Oliva, J., Gazol, A. González de Andrés, E., Montes F., García-Martín, A. y De la Riva, J. 2015. Papel de los factores bióticos y las sequías en el decaimiento del bosque: aportaciones desde la dendroecología. Ecosistemas 24: 15-23. PDF
  8. Sancho-Knapik, D. Peguero-Pina, J.J., Cremer, E., Camarero, J.J., Fernández-Cancio, A., Ibarra, N., Konnert, M. y Gil Pelegrín, E. 2014. Genetic and environmental characterization of Abies alba Mill. populations at its western rear edge. Pirineos 169
  9. Sanz, M.A., Sánchez, J., Camarero, J.J., Peguero-Pina, J.J., Sancho-Knapik, D. y Gil Pelegrín, E. 2014. Change in the terpenoid profile and secondary growth in declining stands of Pinus sylvestris L. under Mediterranean influence as a response to local factors. Pirineos 169 doi: PDF
  10. Camarero, J.J., Vergarechea, M., Sangüesa Barreda, G. y Corrià Ainslie, R. 2014. ¿Condiciona la estructura del monte y el crecimiento previo el decaimiento inducido por la sequía en dos especies de Quercus? Ecosistemas 23: 73-81. PDF
  11. Camarero, J.J., Sangüesa Barreda, G., Alla, A.Q., González de Andrés, E., Maestro Martínez M. y Vicente-Serrano, S.M. 2012. Los precedentes y las respuestas de los árboles a sequías extremas revelan los procesos involucrados en el decaimiento de bosques mediterráneos de coníferas. Ecosistemas 21: 22-30. PDF
  12. Linares, J.C., Pazo-Sarria, R., Taïqui, L., Camarero, J.J., Ochoa, V., Lechuga, V., Seco, J.I., Viñegla, B., Sangüesa, G., Gilarte, P., Merino, J. y Carreira, J.A. 2012. Efectos de las tendencias climáticas y la degradación del hábitat sobre el decaimiento de los cedrales (Cedrus atlantica) del norte de Marruecos. Ecosistemas 21: 7-14. PDF
  13. Sánchez-Salguero, R., Navarro-Cerrillo, R.M., Camarero, J.J., Fernández-Cancio, A., Swetnam, T.W. y Zavala, M.A. 2012. Vulnerabilidad frente a la sequía de repoblaciones de dos especies de pinos en su límite meridional en Europa. Ecosistemas 21: 31-40. PDF
  14. Albuixech, J., Camarero, J.J. y Montserrat-Martí, G. 2012. Dinámica estacional del crecimiento secundario y anatomía del xilema en dos Quercus mediterráneos que coexisten. Forest Systems 21: 9-22. PDF
  15. Camarero, J.J. 2011. Dinámica post-incendio de la vegetación en un valle del Alto Najerilla. Zubía 23: 105-126. PDF
  16. Linares J.C. y Camarero, J. J. 2010. Evaluando los efectos del cambio global sobre el funcionamiento de los abetos ibéricos mediante análisis de isótopos de carbono en los anillos de crecimiento. Ecosistemas, 19/3, Septiembre-Diciembre. PDF
  17. Linares J.C., Camarero, J. J. , Carreira, J.A.,Lamrani-Alaoui, M., Ochoa, V. y García-Ruíz, R. 2009. Patrones espacio-temporales en los pinsapares del macizo Bético-Rifeño. II Reunión del Grupo de Ecología Espacial de la AEET, AEET, Madrid. PDF
  18. Camarero, J. J. 2008. Bosques montanos y subalpinos de Pinus uncinata (*en sustratos yesosos o calcáreos) Hábitat 9430. BASES ECOLÓGICAS PARA LA GESTIÓN DE LOS TIPOS DE HÁBITAT DE INTERÉS COMUNITARIO PRESENTES EN ESPAÑA (DIRECTIVA 92/43/CEE), Dirección General de Conservación de la Naturaleza – AEET, Ministerio de Medio Ambiente, Madrid. PDF
  19. Camarero, J. J., Batllori, E. y Gutiérrez, E. 2008. El límite superior del bosque: interacciones en la frontera del bosque subalpino y los pastos alpinos. Investigación y Ciencia pp. 10-11 (sección Ciencia y Sociedad). PDF
  20. Camarero, J. J. y Gutiérrez, E. 2008. La respuesta del crecimiento de Pinus uncinata al clima en poblaciones relictas del Sistema Ibérico. Zubía 20: 61-96. PDF
  21. Camarero, J. J. y Fortin, M.-J. 2008. Detección cuantitativa de fronteras ecológicas y ecotonos. Introducción al análisis espacial de datos en ecología y ciencias ambientales: métodos y aplicaciones . Maestre, F. T., Escudero, A. y Bonet, A. (eds.). CAM-AEET-URJC, pp. 345-369. PDF
  22. Camarero, J. J. 2006. Dendroecología de Juniperus thurifera en zonas biogeográfica y climáticamente contrastadas. III Coloquio Internacional sobre Sabinares y Enebrales (Género Juniperus): Ecología y Gestión Forestal Sostenible, Soria, Spain. PDF
  23. Camarero, J. J. y Fortin, M.-J. 2006. Detección cuantitativa de fronteras ecológicas y ecotonos. Ecosistemas 2006/3 PDF-www
  24. Sánchez-Miranda, A., Camarero, J.J. y Solla, A. 2006. Efectos del hongo Melampsorella caryophyllacearum Schroet. sobre el crecimiento radial de Abies alba Mill. INIA: Sistemas y Recursos Forestales 15: 171-181. PDF
  25. Camarero, J.J. y Rozas, V. 2006. Técnicas de análisis espacial de patrones de superficies y detección de fronteras aplicadas en ecología forestal. INIA: Sistemas y Recursos Forestales 15: 66-87. PDF
  26. Rozas, V. y Camarero, J.J. 2005. Técnicas de análisis espacial de patrones de puntos aplicadas en ecología forestal. INIA: Sistemas y Recursos Forestales 14: 79-97. PDF
  27. Otero, P., Pintor, M.A., Rodríguez-Rey, L., Camarero, J.J. y Arrechea, E. 2004. El acebo en el Parque Natural del Moncayo: Investigación y gestión forestal. Medio Ambiente-Aragón 18: 18-22. PDF
  28. Camarero, J.J., Lloret, F., Corcuera, L., Peñuelas, J. y Gil-Pelegrín, E. 2004. Cambio global y decaimiento del bosque. Ecología del bosque mediterráneo en un mundo cambiante , Valladares, F. (ed.). Ed. Organismo Autónomo de Parques Nacionales, Ministerio de Medio Ambiente, Madrid, pp. 397-423. PDF PDF-www
  29. Sisó, S., Camarero, J. J., Arrechea, E. y Gil-Pelegrín, E. 2003. Evidencia dendrocronológica sobre el origen natural de Quercus robur L. y de sus híbridos en el Parque del Moncayo. Montes 71: 47-52.
  30. Camarero, J. J., Sisó, S. and Gil-Pelegrín, E. 2003. Fractal dimension does not adequately describe the complexity of leaf margin in seedlings of Quercus species. Anales del Jardín Botánico de Madrid 60: 63-71. PDF
  31. Camarero, J. J., Padró, A., Martín-Bernal, E. y Gil-Pelegrín, E. 2002. Aproximación dendroecológica al decaimiento del abeto (Abies alba Mill.) en el Pirineo aragonés. Montes 70: 26-33. PDF
  32. Camarero, J. J., Martín-Bernal, E. y Gil-Pelegrín, E. 2002. Estudio dendroecológico del ataque de Epinotia subsequana Hw. (Lepidoptera: Tortricidae) sobre un abetar (Abies alba Mill.) del Pirineo aragonés (Diazas, Torla). Inv. Agraria: Sistemas y Recursos Forestales 11: 395-407. PDF
  33. Camarero, J. J. and Gutiérrez, E. 2002. Structure and spatial variability of Pinus uncinata growth forms at two contrasting treeline ecotones in the Spanish Pyrenees. Inv. Agraria: Sistemas y Recursos Forestales 11: 155-169. PDF
  34. Camarero, J. J. 2001. El decaimiento del abeto en los Pirineos. Medio Ambiente-Aragón 4: 18-20. PDF
  35. Camarero, J. J. y Gutiérrez, E. 2001. Spatial variability of tree height at treeline ecotones in the Pyrenees. Orsis 16: 133-144. PDF
  36. Sisó, S., Camarero, J. J. y Gil-Pelegrín, E. 2001. Un nuevo medidor de flujo de alta presión para la estimación de la resistencia hidráulica en brotes completos de plantas leñosas. Inv. Agraria: Sistemas y Recursos Forestales 10: 127-139. PDF
  37. Camarero, J. J. y Gutiérrez, E. 1999. Estructura, patrón espacial y regeneración de una población de Pinus uncinata Ram. en su límite occidental de distribución (Castillo de Vinuesa, Soria-La Rioja). Zubía 17: 99-153. PDF
  38. Camarero, J. J. y Gutiérrez, E. 1999. Estructura de un ecotono bosque subalpino-pastos alpinos (Las Cutas, Ordesa, Pirineos Centrales). Pirineos 153-154: 21-59.  PDF
  39. Camarero, J. J. y Gutiérrez, E. 1999. Estructura espacial de un ecotono bosque subalpino-pastos alpinos en los Estanys de la Pera (Pirineos Orientales) formado por manchas de árboles. Ecología 13: 225-256. PDF
  40. Camarero, J. J. y Gutiérrez, E. 1999. Estructura espacial de un ecotono bosque subalpino-pastos alpinos en el Tessó del Son (Pirineos Centrales) formado por bandas altitudinales de árboles. Ecología 13: 189-224. PDF
  41. Camarero, J. J. y Gutiérrez, E. 1999. Patrón espacial de un ecotono bosque subalpino-pastos alpinos (Las Cutas, Ordesa, Pirineos Centrales. Inv. Agraria: Sistemas y Recursos Forestales 8: 171-205. PDF
  42. Gutiérrez, E., Camarero, J. J. , Tardif, J., Bosch, O. y Ribas, M. 1998. Tendencias recientes del crecimiento y la regeneración en bosques subalpinos del Parque Nacional d’Aigüestortes i Estany de Sant Maurici. Ecología 12: 251-283. PDF
  43. Guerrero-Campo, J., Camarero, J. J. y Gutiérrez, E. 1998. Crecimiento estacional y caída de acículas en Pinus uncinata Ram. y Pinus sylvestris L. Inv. Agraria: Sistemas y Recursos Forestales 7: 155-172. PDF
  44. Camarero, J. J. 1998. Los árboles urbanos también están vivos. Revista de la Asociación Española de Arboricultura 17: 46-46. PDF
  45. Camarero, J. J., Guerrero-Campo, J. y Gutiérrez, E. 1997. Formación del anillo de crecimiento y clima: un ejemplo de modelos para Pinus uncinata Ram. y Pinus sylvestris L. en los Pirineos Centrales. Ecología 11: 235-254. PDF
  46. Camarero, J. J., Guerrero-Campo, J. y Gutiérrez, E. 1996. Fenología del anillo de crecimiento de Pinus uncinata Ram. y Pinus sylvestris L. en un gradiente altitudinal en los Pirineos Centrales. Pirineos 147-148: 3-26. PDF